DNA double strand break repair in Escherichia coli perturbs cell division and chromosome dynamics.

To prevent the transmission of damaged genomic material between generations, cells require a system for accommodating DNA repair within their cell cycles. We have previously shown that Escherichia coli cells subject to a single, repairable site-specific DNA double-strand break (DSB) per DNA replicat...

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Main Authors: Martin A White, Elise Darmon, Manuel A Lopez-Vernaza, David R F Leach
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2020-01-01
Series:PLoS Genetics
Online Access:https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1008473&type=printable
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author Martin A White
Elise Darmon
Manuel A Lopez-Vernaza
David R F Leach
author_facet Martin A White
Elise Darmon
Manuel A Lopez-Vernaza
David R F Leach
author_sort Martin A White
collection DOAJ
description To prevent the transmission of damaged genomic material between generations, cells require a system for accommodating DNA repair within their cell cycles. We have previously shown that Escherichia coli cells subject to a single, repairable site-specific DNA double-strand break (DSB) per DNA replication cycle reach a new average cell length, with a negligible effect on population growth rate. We show here that this new cell size distribution is caused by a DSB repair-dependent delay in completion of cell division. This delay occurs despite unperturbed cell size regulated initiation of both chromosomal DNA replication and cell division. Furthermore, despite DSB repair altering the profile of DNA replication across the genome, the time required to complete chromosomal duplication is invariant. The delay in completion of cell division is accompanied by a DSB repair-dependent delay in individualization of sister nucleoids. We suggest that DSB repair events create inter-sister connections that persist until those chromosomes are separated by a closing septum.
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institution OA Journals
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language English
publishDate 2020-01-01
publisher Public Library of Science (PLoS)
record_format Article
series PLoS Genetics
spelling doaj-art-2cf0688045ee4775adc2114a2d9af9d52025-08-20T02:11:01ZengPublic Library of Science (PLoS)PLoS Genetics1553-73901553-74042020-01-01161e100847310.1371/journal.pgen.1008473DNA double strand break repair in Escherichia coli perturbs cell division and chromosome dynamics.Martin A WhiteElise DarmonManuel A Lopez-VernazaDavid R F LeachTo prevent the transmission of damaged genomic material between generations, cells require a system for accommodating DNA repair within their cell cycles. We have previously shown that Escherichia coli cells subject to a single, repairable site-specific DNA double-strand break (DSB) per DNA replication cycle reach a new average cell length, with a negligible effect on population growth rate. We show here that this new cell size distribution is caused by a DSB repair-dependent delay in completion of cell division. This delay occurs despite unperturbed cell size regulated initiation of both chromosomal DNA replication and cell division. Furthermore, despite DSB repair altering the profile of DNA replication across the genome, the time required to complete chromosomal duplication is invariant. The delay in completion of cell division is accompanied by a DSB repair-dependent delay in individualization of sister nucleoids. We suggest that DSB repair events create inter-sister connections that persist until those chromosomes are separated by a closing septum.https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1008473&type=printable
spellingShingle Martin A White
Elise Darmon
Manuel A Lopez-Vernaza
David R F Leach
DNA double strand break repair in Escherichia coli perturbs cell division and chromosome dynamics.
PLoS Genetics
title DNA double strand break repair in Escherichia coli perturbs cell division and chromosome dynamics.
title_full DNA double strand break repair in Escherichia coli perturbs cell division and chromosome dynamics.
title_fullStr DNA double strand break repair in Escherichia coli perturbs cell division and chromosome dynamics.
title_full_unstemmed DNA double strand break repair in Escherichia coli perturbs cell division and chromosome dynamics.
title_short DNA double strand break repair in Escherichia coli perturbs cell division and chromosome dynamics.
title_sort dna double strand break repair in escherichia coli perturbs cell division and chromosome dynamics
url https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1008473&type=printable
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AT manuelalopezvernaza dnadoublestrandbreakrepairinescherichiacoliperturbscelldivisionandchromosomedynamics
AT davidrfleach dnadoublestrandbreakrepairinescherichiacoliperturbscelldivisionandchromosomedynamics