Veratridine-Induced Oscillations in Nav 1.7 but Not Nav 1.5 Sodium Channels Are Revealed by Membrane Potential Sensitive Dye

Voltage-gated sodium channels (Navs) are critical for membrane potential depolarisation in cells, with especially important roles in neuronal and cardiomyocyte membranes. Their malfunction results in a range of disorders, and they are the target of many widely used drugs. A rapid yet accurate functi...

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Main Authors: Sarah C. R. Lummis, Samantha C. Salvage, Christopher L.-H. Huang, Antony P. Jackson
Format: Article
Language:English
Published: MDPI AG 2025-03-01
Series:Membranes
Subjects:
Online Access:https://www.mdpi.com/2077-0375/15/3/80
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author Sarah C. R. Lummis
Samantha C. Salvage
Christopher L.-H. Huang
Antony P. Jackson
author_facet Sarah C. R. Lummis
Samantha C. Salvage
Christopher L.-H. Huang
Antony P. Jackson
author_sort Sarah C. R. Lummis
collection DOAJ
description Voltage-gated sodium channels (Navs) are critical for membrane potential depolarisation in cells, with especially important roles in neuronal and cardiomyocyte membranes. Their malfunction results in a range of disorders, and they are the target of many widely used drugs. A rapid yet accurate functional assay is therefore desirable both to probe for novel active compounds and to better understand the many different Nav isoforms. Here, we use fluorescence to monitor Nav function: cells expressing either the cardiac Nav 1.5 or pain-associated Nav 1.7 were loaded with fluorescent membrane potential sensitive dye and then stimulated with veratridine. Cells expressing Nav 1.5 show a concentration-dependent slow rise and then a plateau in fluorescence. In contrast, cells expressing Nav 1.7 show a more rapid rise and then unexpected oscillatory behavior. Inhibition by flecainide and mexiletine demonstrates that these oscillations are Nav-dependent. Thus, we show that this fluorescent membrane potential dye can provide useful functional data and that we can readily distinguish between these two Nav isoforms because of the behavior of cells expressing them when activated by veratridine. We consider these distinct behaviors may be due to different interactions of veratridine with the different Nav isoforms, although more studies are needed to understand the mechanism underlying the oscillations.
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spelling doaj-art-2cb02deff7094eeba79081024716d6942025-08-20T01:48:48ZengMDPI AGMembranes2077-03752025-03-011538010.3390/membranes15030080Veratridine-Induced Oscillations in Nav 1.7 but Not Nav 1.5 Sodium Channels Are Revealed by Membrane Potential Sensitive DyeSarah C. R. Lummis0Samantha C. Salvage1Christopher L.-H. Huang2Antony P. Jackson3Department of Biochemistry, University of Cambridge, Tennis Court Road, Cambridge CB2 1QW, UKDepartment of Biochemistry, University of Cambridge, Tennis Court Road, Cambridge CB2 1QW, UKDepartment of Biochemistry, University of Cambridge, Tennis Court Road, Cambridge CB2 1QW, UKDepartment of Biochemistry, University of Cambridge, Tennis Court Road, Cambridge CB2 1QW, UKVoltage-gated sodium channels (Navs) are critical for membrane potential depolarisation in cells, with especially important roles in neuronal and cardiomyocyte membranes. Their malfunction results in a range of disorders, and they are the target of many widely used drugs. A rapid yet accurate functional assay is therefore desirable both to probe for novel active compounds and to better understand the many different Nav isoforms. Here, we use fluorescence to monitor Nav function: cells expressing either the cardiac Nav 1.5 or pain-associated Nav 1.7 were loaded with fluorescent membrane potential sensitive dye and then stimulated with veratridine. Cells expressing Nav 1.5 show a concentration-dependent slow rise and then a plateau in fluorescence. In contrast, cells expressing Nav 1.7 show a more rapid rise and then unexpected oscillatory behavior. Inhibition by flecainide and mexiletine demonstrates that these oscillations are Nav-dependent. Thus, we show that this fluorescent membrane potential dye can provide useful functional data and that we can readily distinguish between these two Nav isoforms because of the behavior of cells expressing them when activated by veratridine. We consider these distinct behaviors may be due to different interactions of veratridine with the different Nav isoforms, although more studies are needed to understand the mechanism underlying the oscillations.https://www.mdpi.com/2077-0375/15/3/80oscillationsdepolarizationFlexstationvoltage-gated sodium channelvoltage-sensitive dye
spellingShingle Sarah C. R. Lummis
Samantha C. Salvage
Christopher L.-H. Huang
Antony P. Jackson
Veratridine-Induced Oscillations in Nav 1.7 but Not Nav 1.5 Sodium Channels Are Revealed by Membrane Potential Sensitive Dye
Membranes
oscillations
depolarization
Flexstation
voltage-gated sodium channel
voltage-sensitive dye
title Veratridine-Induced Oscillations in Nav 1.7 but Not Nav 1.5 Sodium Channels Are Revealed by Membrane Potential Sensitive Dye
title_full Veratridine-Induced Oscillations in Nav 1.7 but Not Nav 1.5 Sodium Channels Are Revealed by Membrane Potential Sensitive Dye
title_fullStr Veratridine-Induced Oscillations in Nav 1.7 but Not Nav 1.5 Sodium Channels Are Revealed by Membrane Potential Sensitive Dye
title_full_unstemmed Veratridine-Induced Oscillations in Nav 1.7 but Not Nav 1.5 Sodium Channels Are Revealed by Membrane Potential Sensitive Dye
title_short Veratridine-Induced Oscillations in Nav 1.7 but Not Nav 1.5 Sodium Channels Are Revealed by Membrane Potential Sensitive Dye
title_sort veratridine induced oscillations in nav 1 7 but not nav 1 5 sodium channels are revealed by membrane potential sensitive dye
topic oscillations
depolarization
Flexstation
voltage-gated sodium channel
voltage-sensitive dye
url https://www.mdpi.com/2077-0375/15/3/80
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