Accounting for imperfect detection when estimating species‐area relationships and beta‐diversity

Abstract Ecologists have historically quantified fundamental biodiversity patterns, including species‐area relationships (SARs) and beta diversity, using observed species counts. However, imperfect detection may often bias derived community metrics and subsequent community models. Although several s...

Full description

Saved in:
Bibliographic Details
Main Authors: Ciar D. Noble, Carlos A. Peres, James J. Gilroy
Format: Article
Language:English
Published: Wiley 2024-07-01
Series:Ecology and Evolution
Subjects:
Online Access:https://doi.org/10.1002/ece3.70017
Tags: Add Tag
No Tags, Be the first to tag this record!
_version_ 1849714086203359232
author Ciar D. Noble
Carlos A. Peres
James J. Gilroy
author_facet Ciar D. Noble
Carlos A. Peres
James J. Gilroy
author_sort Ciar D. Noble
collection DOAJ
description Abstract Ecologists have historically quantified fundamental biodiversity patterns, including species‐area relationships (SARs) and beta diversity, using observed species counts. However, imperfect detection may often bias derived community metrics and subsequent community models. Although several statistical methods claim to correct for imperfect detection, their performance in species‐area and β‐diversity research remains unproven. We examine inaccuracies in the estimation of SARs and β‐diversity parameters that emerge from imperfect detection, and whether such errors can be mitigated using a non‐parametric diversity estimator (iNEXT.3D) and Multi‐Species Occupancy Models (MSOMs). We simulated 28,350 sampling regimes of 2835 fragmented communities, varying the mean and standard deviation of species detection probabilities, and the number of sampling repetitions. We then quantified the bias, accuracy, and precision of derived estimates of model coefficients for SARs and the effects of patch area on β‐diversity (pairwise Sørensen similarity). Imperfect detection biased estimates of all evaluated parameters, particularly when mean detection probabilities were low, and there were few sampling repetitions. Observed counts consistently underestimated species richness and SAR z‐values, and overestimated SAR c‐values; iNEXT.3D and MSOMs only partially resolved these biases. iNEXT.3D provided the best estimates of SAR z‐values, although MSOM estimates were generally comparable. All three methods accurately estimated pairwise Sørensen similarity in most circumstances, but only MSOMs provided unbiased estimates of the coefficients of models examining covariate effects on β‐diversity. Even when using iNEXT.3D or MSOMs, imperfect detection consistently caused biases in SAR coefficient estimates, calling into question the robustness of previous SAR studies. Furthermore, the inability of observed counts and iNEXT.3D to estimate β‐diversity model coefficients resulted from a systematic, area‐related bias in Sørensen similarity estimates. Importantly, MSOMs corrected for these biases in β‐diversity assessments, even in suboptimal scenarios. Nonetheless, as estimator performance consistently improved with increasing sampling repetitions, the importance of appropriate sampling effort cannot be understated.
format Article
id doaj-art-2ae9502b7bce42b4be920bf60943273a
institution DOAJ
issn 2045-7758
language English
publishDate 2024-07-01
publisher Wiley
record_format Article
series Ecology and Evolution
spelling doaj-art-2ae9502b7bce42b4be920bf60943273a2025-08-20T03:13:48ZengWileyEcology and Evolution2045-77582024-07-01147n/an/a10.1002/ece3.70017Accounting for imperfect detection when estimating species‐area relationships and beta‐diversityCiar D. Noble0Carlos A. Peres1James J. Gilroy2School of Environmental Sciences University of East Anglia Norwich, Norfolk UKSchool of Environmental Sciences University of East Anglia Norwich, Norfolk UKSchool of Environmental Sciences University of East Anglia Norwich, Norfolk UKAbstract Ecologists have historically quantified fundamental biodiversity patterns, including species‐area relationships (SARs) and beta diversity, using observed species counts. However, imperfect detection may often bias derived community metrics and subsequent community models. Although several statistical methods claim to correct for imperfect detection, their performance in species‐area and β‐diversity research remains unproven. We examine inaccuracies in the estimation of SARs and β‐diversity parameters that emerge from imperfect detection, and whether such errors can be mitigated using a non‐parametric diversity estimator (iNEXT.3D) and Multi‐Species Occupancy Models (MSOMs). We simulated 28,350 sampling regimes of 2835 fragmented communities, varying the mean and standard deviation of species detection probabilities, and the number of sampling repetitions. We then quantified the bias, accuracy, and precision of derived estimates of model coefficients for SARs and the effects of patch area on β‐diversity (pairwise Sørensen similarity). Imperfect detection biased estimates of all evaluated parameters, particularly when mean detection probabilities were low, and there were few sampling repetitions. Observed counts consistently underestimated species richness and SAR z‐values, and overestimated SAR c‐values; iNEXT.3D and MSOMs only partially resolved these biases. iNEXT.3D provided the best estimates of SAR z‐values, although MSOM estimates were generally comparable. All three methods accurately estimated pairwise Sørensen similarity in most circumstances, but only MSOMs provided unbiased estimates of the coefficients of models examining covariate effects on β‐diversity. Even when using iNEXT.3D or MSOMs, imperfect detection consistently caused biases in SAR coefficient estimates, calling into question the robustness of previous SAR studies. Furthermore, the inability of observed counts and iNEXT.3D to estimate β‐diversity model coefficients resulted from a systematic, area‐related bias in Sørensen similarity estimates. Importantly, MSOMs corrected for these biases in β‐diversity assessments, even in suboptimal scenarios. Nonetheless, as estimator performance consistently improved with increasing sampling repetitions, the importance of appropriate sampling effort cannot be understated.https://doi.org/10.1002/ece3.70017Chao estimatorcommunity simulationhabitat fragmentationiNEXT.3Dmulti‐species occupancy modelspecies richness
spellingShingle Ciar D. Noble
Carlos A. Peres
James J. Gilroy
Accounting for imperfect detection when estimating species‐area relationships and beta‐diversity
Ecology and Evolution
Chao estimator
community simulation
habitat fragmentation
iNEXT.3D
multi‐species occupancy model
species richness
title Accounting for imperfect detection when estimating species‐area relationships and beta‐diversity
title_full Accounting for imperfect detection when estimating species‐area relationships and beta‐diversity
title_fullStr Accounting for imperfect detection when estimating species‐area relationships and beta‐diversity
title_full_unstemmed Accounting for imperfect detection when estimating species‐area relationships and beta‐diversity
title_short Accounting for imperfect detection when estimating species‐area relationships and beta‐diversity
title_sort accounting for imperfect detection when estimating species area relationships and beta diversity
topic Chao estimator
community simulation
habitat fragmentation
iNEXT.3D
multi‐species occupancy model
species richness
url https://doi.org/10.1002/ece3.70017
work_keys_str_mv AT ciardnoble accountingforimperfectdetectionwhenestimatingspeciesarearelationshipsandbetadiversity
AT carlosaperes accountingforimperfectdetectionwhenestimatingspeciesarearelationshipsandbetadiversity
AT jamesjgilroy accountingforimperfectdetectionwhenestimatingspeciesarearelationshipsandbetadiversity