Phosphatidylserine synthase regulates cellular homeostasis through distinct metabolic mechanisms.

Phosphatidylserine (PS), synthesized in the endoplasmic reticulum (ER) by phosphatidylserine synthase (PSS), is transported to the plasma membrane (PM) and mitochondria through distinct routes. The in vivo functions of PS at different subcellular locations and the coordination between different PS t...

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Main Authors: Xiao Yang, Jingjing Liang, Long Ding, Xia Li, Sin-Man Lam, Guanghou Shui, Mei Ding, Xun Huang
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2019-12-01
Series:PLoS Genetics
Online Access:https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1008548&type=printable
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author Xiao Yang
Jingjing Liang
Long Ding
Xia Li
Sin-Man Lam
Guanghou Shui
Mei Ding
Xun Huang
author_facet Xiao Yang
Jingjing Liang
Long Ding
Xia Li
Sin-Man Lam
Guanghou Shui
Mei Ding
Xun Huang
author_sort Xiao Yang
collection DOAJ
description Phosphatidylserine (PS), synthesized in the endoplasmic reticulum (ER) by phosphatidylserine synthase (PSS), is transported to the plasma membrane (PM) and mitochondria through distinct routes. The in vivo functions of PS at different subcellular locations and the coordination between different PS transport routes are not fully understood. Here, we report that Drosophila PSS regulates cell growth, lipid storage and mitochondrial function. In pss RNAi, reduced PS depletes plasma membrane Akt, contributing to cell growth defects; the metabolic shift from phospholipid synthesis to neutral lipid synthesis results in ectopic lipid accumulation; and the reduction of mitochondrial PS impairs mitochondrial protein import and mitochondrial integrity. Importantly, reducing PS transport from the ER to PM by loss of PI4KIIIα partially rescues the mitochondrial defects of pss RNAi. Together, our results uncover a balance between different PS transport routes and reveal that PSS regulates cellular homeostasis through distinct metabolic mechanisms.
format Article
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institution OA Journals
issn 1553-7390
1553-7404
language English
publishDate 2019-12-01
publisher Public Library of Science (PLoS)
record_format Article
series PLoS Genetics
spelling doaj-art-283ae42a327b48048daf2a2175f0c7702025-08-20T02:11:01ZengPublic Library of Science (PLoS)PLoS Genetics1553-73901553-74042019-12-011512e100854810.1371/journal.pgen.1008548Phosphatidylserine synthase regulates cellular homeostasis through distinct metabolic mechanisms.Xiao YangJingjing LiangLong DingXia LiSin-Man LamGuanghou ShuiMei DingXun HuangPhosphatidylserine (PS), synthesized in the endoplasmic reticulum (ER) by phosphatidylserine synthase (PSS), is transported to the plasma membrane (PM) and mitochondria through distinct routes. The in vivo functions of PS at different subcellular locations and the coordination between different PS transport routes are not fully understood. Here, we report that Drosophila PSS regulates cell growth, lipid storage and mitochondrial function. In pss RNAi, reduced PS depletes plasma membrane Akt, contributing to cell growth defects; the metabolic shift from phospholipid synthesis to neutral lipid synthesis results in ectopic lipid accumulation; and the reduction of mitochondrial PS impairs mitochondrial protein import and mitochondrial integrity. Importantly, reducing PS transport from the ER to PM by loss of PI4KIIIα partially rescues the mitochondrial defects of pss RNAi. Together, our results uncover a balance between different PS transport routes and reveal that PSS regulates cellular homeostasis through distinct metabolic mechanisms.https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1008548&type=printable
spellingShingle Xiao Yang
Jingjing Liang
Long Ding
Xia Li
Sin-Man Lam
Guanghou Shui
Mei Ding
Xun Huang
Phosphatidylserine synthase regulates cellular homeostasis through distinct metabolic mechanisms.
PLoS Genetics
title Phosphatidylserine synthase regulates cellular homeostasis through distinct metabolic mechanisms.
title_full Phosphatidylserine synthase regulates cellular homeostasis through distinct metabolic mechanisms.
title_fullStr Phosphatidylserine synthase regulates cellular homeostasis through distinct metabolic mechanisms.
title_full_unstemmed Phosphatidylserine synthase regulates cellular homeostasis through distinct metabolic mechanisms.
title_short Phosphatidylserine synthase regulates cellular homeostasis through distinct metabolic mechanisms.
title_sort phosphatidylserine synthase regulates cellular homeostasis through distinct metabolic mechanisms
url https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1008548&type=printable
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