Fossil constraints on the origin and evolution of Platyhelminthes are surprisingly concordant with modern molecular phylogenies

ABSTRACT Trace fossils preserved with fossil worm-shaped remains suggest the presence of free-living flatworms during the Ordovician at the latest and their occurrence in terrestrial environments during the Permian. The presence of hooks associated with acanthodian and placoderm fishes indicates the...

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Main Authors: Kenneth De Baets, Paula Dentzien-Dias, John Warren Huntley, Maarten P.M. Vanhove, Weronika Łaska, Aleksandra Skawina, Niels W.L. Van Steenkiste, Karina Vanadzina
Format: Article
Language:English
Published: Sociedade Brasileira de Zoologia 2024-11-01
Series:Zoologia (Curitiba)
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Online Access:http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1984-46702024000100708&lng=en&tlng=en
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author Kenneth De Baets
Paula Dentzien-Dias
John Warren Huntley
Maarten P.M. Vanhove
Weronika Łaska
Aleksandra Skawina
Niels W.L. Van Steenkiste
Karina Vanadzina
author_facet Kenneth De Baets
Paula Dentzien-Dias
John Warren Huntley
Maarten P.M. Vanhove
Weronika Łaska
Aleksandra Skawina
Niels W.L. Van Steenkiste
Karina Vanadzina
author_sort Kenneth De Baets
collection DOAJ
description ABSTRACT Trace fossils preserved with fossil worm-shaped remains suggest the presence of free-living flatworms during the Ordovician at the latest and their occurrence in terrestrial environments during the Permian. The presence of hooks associated with acanthodian and placoderm fishes indicates the existence of parasitic monopisthocotyleans, with a simple life cycle, during the Devonian. The presence of eggs in shark coprolites suggests the occurrence of eucestode tapeworms, with complex life cycles, during the Permian, possibly even earlier in the Carboniferous. Fossil evidence for trematode flatworms, also with complex life cycles, is more recent, including diverse findings associated with bivalves, lizards, and coprolites of archosaurs in terrestrial environments between 126 and 76 Ma in the Cretaceous. Convincing evidence for gymnophallid trematodes in marine environments appears in the Eocene, with an earlier occurrence in Cretaceous freshwater environments. This chronological pattern of first appearance (Turbellarians > Monopisthocotylea > Cestoda > Trematoda) is surprisingly concordant with some recent molecular phylogenetic analyses. Further evidence to test these hypotheses could be obtained by conducting systematic screenings for resistant remains of platyhelminths such as hooks and eggs as well as characteristic traces such as trails or shell concretions preserved with their producers. Additional study and scrutiny are particularly needed for trace fossils attributed to free-living flatworms that are not associated with their producers. We make recommendations on how different constraints on flatworm evolution can be interpreted and used in future studies.
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spelling doaj-art-26bc4e3e773e4047a379b37539e1d45c2025-08-20T02:36:45ZengSociedade Brasileira de ZoologiaZoologia (Curitiba)1984-46892024-11-014110.1590/s1984-4689.v41.e24002Fossil constraints on the origin and evolution of Platyhelminthes are surprisingly concordant with modern molecular phylogeniesKenneth De Baetshttps://orcid.org/0000-0002-1651-321XPaula Dentzien-Diashttps://orcid.org/0000-0001-8708-6799John Warren Huntleyhttps://orcid.org/0000-0001-9955-8853Maarten P.M. Vanhovehttps://orcid.org/0000-0003-3100-7566Weronika Łaskahttps://orcid.org/0000-0001-5034-5240Aleksandra Skawinahttps://orcid.org/0000-0002-8287-1568Niels W.L. Van Steenkistehttps://orcid.org/0000-0002-2676-7862Karina Vanadzinahttps://orcid.org/0000-0002-7952-2658ABSTRACT Trace fossils preserved with fossil worm-shaped remains suggest the presence of free-living flatworms during the Ordovician at the latest and their occurrence in terrestrial environments during the Permian. The presence of hooks associated with acanthodian and placoderm fishes indicates the existence of parasitic monopisthocotyleans, with a simple life cycle, during the Devonian. The presence of eggs in shark coprolites suggests the occurrence of eucestode tapeworms, with complex life cycles, during the Permian, possibly even earlier in the Carboniferous. Fossil evidence for trematode flatworms, also with complex life cycles, is more recent, including diverse findings associated with bivalves, lizards, and coprolites of archosaurs in terrestrial environments between 126 and 76 Ma in the Cretaceous. Convincing evidence for gymnophallid trematodes in marine environments appears in the Eocene, with an earlier occurrence in Cretaceous freshwater environments. This chronological pattern of first appearance (Turbellarians > Monopisthocotylea > Cestoda > Trematoda) is surprisingly concordant with some recent molecular phylogenetic analyses. Further evidence to test these hypotheses could be obtained by conducting systematic screenings for resistant remains of platyhelminths such as hooks and eggs as well as characteristic traces such as trails or shell concretions preserved with their producers. Additional study and scrutiny are particularly needed for trace fossils attributed to free-living flatworms that are not associated with their producers. We make recommendations on how different constraints on flatworm evolution can be interpreted and used in future studies.http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1984-46702024000100708&lng=en&tlng=enCophylogenyflatwormsfossil recordmacroevolution
spellingShingle Kenneth De Baets
Paula Dentzien-Dias
John Warren Huntley
Maarten P.M. Vanhove
Weronika Łaska
Aleksandra Skawina
Niels W.L. Van Steenkiste
Karina Vanadzina
Fossil constraints on the origin and evolution of Platyhelminthes are surprisingly concordant with modern molecular phylogenies
Zoologia (Curitiba)
Cophylogeny
flatworms
fossil record
macroevolution
title Fossil constraints on the origin and evolution of Platyhelminthes are surprisingly concordant with modern molecular phylogenies
title_full Fossil constraints on the origin and evolution of Platyhelminthes are surprisingly concordant with modern molecular phylogenies
title_fullStr Fossil constraints on the origin and evolution of Platyhelminthes are surprisingly concordant with modern molecular phylogenies
title_full_unstemmed Fossil constraints on the origin and evolution of Platyhelminthes are surprisingly concordant with modern molecular phylogenies
title_short Fossil constraints on the origin and evolution of Platyhelminthes are surprisingly concordant with modern molecular phylogenies
title_sort fossil constraints on the origin and evolution of platyhelminthes are surprisingly concordant with modern molecular phylogenies
topic Cophylogeny
flatworms
fossil record
macroevolution
url http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1984-46702024000100708&lng=en&tlng=en
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