Quorum Sensing Coordinates Carbon and Nitrogen Metabolism to Optimize Public Goods Production in Pseudomonas fluorescens 2P24

Abstract The coordination of public and private goods production is essential for bacterial adaptation to environmental changes. Quorum sensing (QS) regulates this balance by mediating the trade‐off between the communal benefits of “public goods,” such as siderophores and antibiotics, and the indivi...

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Main Authors: Jie Li, Mengxue Nie, Hongguang Ma, Xuanying Tao, Yanxia Sun, Xinyue Tu, Pingping Zhang, Li‐Qun Zhang, Rong Jia, Yong‐Xing He, Nannan Zhang, Honghua Ge
Format: Article
Language:English
Published: Wiley 2025-03-01
Series:Advanced Science
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Online Access:https://doi.org/10.1002/advs.202412224
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author Jie Li
Mengxue Nie
Hongguang Ma
Xuanying Tao
Yanxia Sun
Xinyue Tu
Pingping Zhang
Li‐Qun Zhang
Rong Jia
Yong‐Xing He
Nannan Zhang
Honghua Ge
author_facet Jie Li
Mengxue Nie
Hongguang Ma
Xuanying Tao
Yanxia Sun
Xinyue Tu
Pingping Zhang
Li‐Qun Zhang
Rong Jia
Yong‐Xing He
Nannan Zhang
Honghua Ge
author_sort Jie Li
collection DOAJ
description Abstract The coordination of public and private goods production is essential for bacterial adaptation to environmental changes. Quorum sensing (QS) regulates this balance by mediating the trade‐off between the communal benefits of “public goods,” such as siderophores and antibiotics, and the individual metabolic needs fulfilled by “private goods,” such as intracellular metabolites utilized for growth and survival. Pseudomonas fluorescens 2P24 harbors a LasI/LasR‐type QS system, MupI/MupR, which regulates mupirocin production through signaling molecules. This study explores how QS coordinates carbon and nitrogen metabolism to optimize the production of key secondary metabolites, including 2,4‐diacetylphloroglucinol (2,4‐DAPG), mupirocin, and siderophores, which serve as public goods. Loss of QS disrupts this balance by enhancing the Krebs cycle, denitrification, pyruvate anaplerosis, and ammonium assimilation, lead to halted 2,4‐DAPG and mupirocin synthesis and increased siderophore production. In the absence of QS, elevated siderophore production compensates for iron acquisition, ensuring rapid cellular growth. Under nutrient‐limited or high cell density conditions, MupR regulates carbon and nitrogen fluxes to sustain public goods production. These findings highlight QS as a key environmental sensor that fine‐tunes resource allocation, bacterial fitness, and adaptation to ecological and nutritional conditions, suggesting the potential for QS‐targeted approaches to enhance antibiotic production and agricultural sustainability.
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spelling doaj-art-25f340fdf8684c47987c97e9b75c0cdf2025-08-20T02:10:46ZengWileyAdvanced Science2198-38442025-03-011212n/an/a10.1002/advs.202412224Quorum Sensing Coordinates Carbon and Nitrogen Metabolism to Optimize Public Goods Production in Pseudomonas fluorescens 2P24Jie Li0Mengxue Nie1Hongguang Ma2Xuanying Tao3Yanxia Sun4Xinyue Tu5Pingping Zhang6Li‐Qun Zhang7Rong Jia8Yong‐Xing He9Nannan Zhang10Honghua Ge11School of Life Sciences Anhui University Hefei 230601 ChinaSchool of Life Sciences Anhui University Hefei 230601 ChinaSchool of Life Sciences Anhui University Hefei 230601 ChinaSchool of Life Sciences Anhui University Hefei 230601 ChinaSchool of Life Sciences Anhui University Hefei 230601 ChinaSchool of Life Sciences Anhui University Hefei 230601 ChinaSchool of Life Sciences Anhui University Hefei 230601 ChinaCollege of Plant Protection China Agricultural University Beijing 100083 ChinaSchool of Life Sciences Anhui University Hefei 230601 ChinaMinistry of Education Key Laboratory of Cell Activities and Stress Adaptations School of Life Sciences Lanzhou University Lanzhou 730000 ChinaSchool of Life Sciences Anhui University Hefei 230601 ChinaInstitute of Health Sciences and Technology Institutes of Physical Science and Information Technology Anhui University Hefei 230601 ChinaAbstract The coordination of public and private goods production is essential for bacterial adaptation to environmental changes. Quorum sensing (QS) regulates this balance by mediating the trade‐off between the communal benefits of “public goods,” such as siderophores and antibiotics, and the individual metabolic needs fulfilled by “private goods,” such as intracellular metabolites utilized for growth and survival. Pseudomonas fluorescens 2P24 harbors a LasI/LasR‐type QS system, MupI/MupR, which regulates mupirocin production through signaling molecules. This study explores how QS coordinates carbon and nitrogen metabolism to optimize the production of key secondary metabolites, including 2,4‐diacetylphloroglucinol (2,4‐DAPG), mupirocin, and siderophores, which serve as public goods. Loss of QS disrupts this balance by enhancing the Krebs cycle, denitrification, pyruvate anaplerosis, and ammonium assimilation, lead to halted 2,4‐DAPG and mupirocin synthesis and increased siderophore production. In the absence of QS, elevated siderophore production compensates for iron acquisition, ensuring rapid cellular growth. Under nutrient‐limited or high cell density conditions, MupR regulates carbon and nitrogen fluxes to sustain public goods production. These findings highlight QS as a key environmental sensor that fine‐tunes resource allocation, bacterial fitness, and adaptation to ecological and nutritional conditions, suggesting the potential for QS‐targeted approaches to enhance antibiotic production and agricultural sustainability.https://doi.org/10.1002/advs.202412224carbon and nitrogen metabolismgene transcriptionpublic goodsquorum sensingtranscription factor
spellingShingle Jie Li
Mengxue Nie
Hongguang Ma
Xuanying Tao
Yanxia Sun
Xinyue Tu
Pingping Zhang
Li‐Qun Zhang
Rong Jia
Yong‐Xing He
Nannan Zhang
Honghua Ge
Quorum Sensing Coordinates Carbon and Nitrogen Metabolism to Optimize Public Goods Production in Pseudomonas fluorescens 2P24
Advanced Science
carbon and nitrogen metabolism
gene transcription
public goods
quorum sensing
transcription factor
title Quorum Sensing Coordinates Carbon and Nitrogen Metabolism to Optimize Public Goods Production in Pseudomonas fluorescens 2P24
title_full Quorum Sensing Coordinates Carbon and Nitrogen Metabolism to Optimize Public Goods Production in Pseudomonas fluorescens 2P24
title_fullStr Quorum Sensing Coordinates Carbon and Nitrogen Metabolism to Optimize Public Goods Production in Pseudomonas fluorescens 2P24
title_full_unstemmed Quorum Sensing Coordinates Carbon and Nitrogen Metabolism to Optimize Public Goods Production in Pseudomonas fluorescens 2P24
title_short Quorum Sensing Coordinates Carbon and Nitrogen Metabolism to Optimize Public Goods Production in Pseudomonas fluorescens 2P24
title_sort quorum sensing coordinates carbon and nitrogen metabolism to optimize public goods production in pseudomonas fluorescens 2p24
topic carbon and nitrogen metabolism
gene transcription
public goods
quorum sensing
transcription factor
url https://doi.org/10.1002/advs.202412224
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