Role of DNA methylation in expression control of the IKZF3-GSDMA region in human epithelial cells.

Chromosomal region 17q12-q21 is associated with asthma and harbors regulatory polymorphisms that influence expression levels of all five protein-coding genes in the region: IKAROS family zinc finger 3 (Aiolos) (IKZF3), zona pellucida binding protein 2 (ZPBP2), ORMDL sphingolipid biosynthesis regulat...

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Main Authors: Sanny Moussette, Abeer Al Tuwaijri, Hamid-Reza Kohan-Ghadr, Samar Elzein, Raquel Farias, Julie Bérubé, Bianca Ho, Catherine Laprise, Cynthia G Goodyer, Simon Rousseau, Anna K Naumova
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2017-01-01
Series:PLoS ONE
Online Access:https://journals.plos.org/plosone/article/file?id=10.1371/journal.pone.0172707&type=printable
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author Sanny Moussette
Abeer Al Tuwaijri
Hamid-Reza Kohan-Ghadr
Samar Elzein
Raquel Farias
Julie Bérubé
Bianca Ho
Catherine Laprise
Cynthia G Goodyer
Simon Rousseau
Anna K Naumova
author_facet Sanny Moussette
Abeer Al Tuwaijri
Hamid-Reza Kohan-Ghadr
Samar Elzein
Raquel Farias
Julie Bérubé
Bianca Ho
Catherine Laprise
Cynthia G Goodyer
Simon Rousseau
Anna K Naumova
author_sort Sanny Moussette
collection DOAJ
description Chromosomal region 17q12-q21 is associated with asthma and harbors regulatory polymorphisms that influence expression levels of all five protein-coding genes in the region: IKAROS family zinc finger 3 (Aiolos) (IKZF3), zona pellucida binding protein 2 (ZPBP2), ORMDL sphingolipid biosynthesis regulator 3 (ORMDL3), and gasdermins A and B (GSDMA, GSDMB). Furthermore, DNA methylation in this region has been implicated as a potential modifier of the genetic risk of asthma development. To further characterize the effect of DNA methylation, we examined the impact of treatment with DNA methyltransferase inhibitor 5-aza-2'-deoxycytidine (5-aza-dC) that causes DNA demethylation, on expression and promoter methylation of the five 17q12-q21 genes in the human airway epithelium cell line NuLi-1, embryonic kidney epithelium cell line 293T and human adenocarcinoma cell line MCF-7. 5-aza-dC treatment led to upregulation of expression of GSDMA in all three cell lines. ZPBP2 was upregulated in NuLi-1, but remained repressed in 293T and MCF-7 cells, whereas ORMDL3 was upregulated in 293T and MCF-7 cells, but not NuLi-1. Upregulation of ZPBP2 and GSDMA was accompanied by a decrease in promoter methylation. Moreover, 5-aza-dC treatment modified allelic expression of ZPBP2 and ORMDL3 suggesting that different alleles may respond differently to treatment. We also identified a polymorphic CTCF-binding site in intron 1 of ORMDL3 carrying a CG SNP rs4065275 and determined its methylation level. The site's methylation was unaffected by 5-aza-dC treatment in NuLi-1 cells. We conclude that modest changes (8-13%) in promoter methylation levels of ZPBP2 and GSDMA may cause substantial changes in RNA levels and that allelic expression of ZPBP2 and ORMDL3 is mediated by DNA methylation.
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spelling doaj-art-233d149f5aaf4ebd8b8ca868f729fed32025-08-20T03:24:29ZengPublic Library of Science (PLoS)PLoS ONE1932-62032017-01-01122e017270710.1371/journal.pone.0172707Role of DNA methylation in expression control of the IKZF3-GSDMA region in human epithelial cells.Sanny MoussetteAbeer Al TuwaijriHamid-Reza Kohan-GhadrSamar ElzeinRaquel FariasJulie BérubéBianca HoCatherine LapriseCynthia G GoodyerSimon RousseauAnna K NaumovaChromosomal region 17q12-q21 is associated with asthma and harbors regulatory polymorphisms that influence expression levels of all five protein-coding genes in the region: IKAROS family zinc finger 3 (Aiolos) (IKZF3), zona pellucida binding protein 2 (ZPBP2), ORMDL sphingolipid biosynthesis regulator 3 (ORMDL3), and gasdermins A and B (GSDMA, GSDMB). Furthermore, DNA methylation in this region has been implicated as a potential modifier of the genetic risk of asthma development. To further characterize the effect of DNA methylation, we examined the impact of treatment with DNA methyltransferase inhibitor 5-aza-2'-deoxycytidine (5-aza-dC) that causes DNA demethylation, on expression and promoter methylation of the five 17q12-q21 genes in the human airway epithelium cell line NuLi-1, embryonic kidney epithelium cell line 293T and human adenocarcinoma cell line MCF-7. 5-aza-dC treatment led to upregulation of expression of GSDMA in all three cell lines. ZPBP2 was upregulated in NuLi-1, but remained repressed in 293T and MCF-7 cells, whereas ORMDL3 was upregulated in 293T and MCF-7 cells, but not NuLi-1. Upregulation of ZPBP2 and GSDMA was accompanied by a decrease in promoter methylation. Moreover, 5-aza-dC treatment modified allelic expression of ZPBP2 and ORMDL3 suggesting that different alleles may respond differently to treatment. We also identified a polymorphic CTCF-binding site in intron 1 of ORMDL3 carrying a CG SNP rs4065275 and determined its methylation level. The site's methylation was unaffected by 5-aza-dC treatment in NuLi-1 cells. We conclude that modest changes (8-13%) in promoter methylation levels of ZPBP2 and GSDMA may cause substantial changes in RNA levels and that allelic expression of ZPBP2 and ORMDL3 is mediated by DNA methylation.https://journals.plos.org/plosone/article/file?id=10.1371/journal.pone.0172707&type=printable
spellingShingle Sanny Moussette
Abeer Al Tuwaijri
Hamid-Reza Kohan-Ghadr
Samar Elzein
Raquel Farias
Julie Bérubé
Bianca Ho
Catherine Laprise
Cynthia G Goodyer
Simon Rousseau
Anna K Naumova
Role of DNA methylation in expression control of the IKZF3-GSDMA region in human epithelial cells.
PLoS ONE
title Role of DNA methylation in expression control of the IKZF3-GSDMA region in human epithelial cells.
title_full Role of DNA methylation in expression control of the IKZF3-GSDMA region in human epithelial cells.
title_fullStr Role of DNA methylation in expression control of the IKZF3-GSDMA region in human epithelial cells.
title_full_unstemmed Role of DNA methylation in expression control of the IKZF3-GSDMA region in human epithelial cells.
title_short Role of DNA methylation in expression control of the IKZF3-GSDMA region in human epithelial cells.
title_sort role of dna methylation in expression control of the ikzf3 gsdma region in human epithelial cells
url https://journals.plos.org/plosone/article/file?id=10.1371/journal.pone.0172707&type=printable
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