SLC25A46 is required for mitochondrial lipid homeostasis and cristae maintenance and is responsible for Leigh syndrome

Abstract Mitochondria form a dynamic network that responds to physiological signals and metabolic stresses by altering the balance between fusion and fission. Mitochondrial fusion is orchestrated by conserved GTPases MFN1/2 and OPA1, a process coordinated in yeast by Ugo1, a mitochondrial metabolite...

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Main Authors: Alexandre Janer, Julien Prudent, Vincent Paupe, Somayyeh Fahiminiya, Jacek Majewski, Nicolas Sgarioto, Christine Des Rosiers, Anik Forest, Zhen‐Yuan Lin, Anne‐Claude Gingras, Grant Mitchell, Heidi M McBride, Eric A Shoubridge
Format: Article
Language:English
Published: Springer Nature 2016-07-01
Series:EMBO Molecular Medicine
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Online Access:https://doi.org/10.15252/emmm.201506159
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author Alexandre Janer
Julien Prudent
Vincent Paupe
Somayyeh Fahiminiya
Jacek Majewski
Nicolas Sgarioto
Christine Des Rosiers
Anik Forest
Zhen‐Yuan Lin
Anne‐Claude Gingras
Grant Mitchell
Heidi M McBride
Eric A Shoubridge
author_facet Alexandre Janer
Julien Prudent
Vincent Paupe
Somayyeh Fahiminiya
Jacek Majewski
Nicolas Sgarioto
Christine Des Rosiers
Anik Forest
Zhen‐Yuan Lin
Anne‐Claude Gingras
Grant Mitchell
Heidi M McBride
Eric A Shoubridge
author_sort Alexandre Janer
collection DOAJ
description Abstract Mitochondria form a dynamic network that responds to physiological signals and metabolic stresses by altering the balance between fusion and fission. Mitochondrial fusion is orchestrated by conserved GTPases MFN1/2 and OPA1, a process coordinated in yeast by Ugo1, a mitochondrial metabolite carrier family protein. We uncovered a homozygous missense mutation in SLC25A46, the mammalian orthologue of Ugo1, in a subject with Leigh syndrome. SLC25A46 is an integral outer membrane protein that interacts with MFN2, OPA1, and the mitochondrial contact site and cristae organizing system (MICOS) complex. The subject mutation destabilizes the protein, leading to mitochondrial hyperfusion, alterations in endoplasmic reticulum (ER) morphology, impaired cellular respiration, and premature cellular senescence. The MICOS complex is disrupted in subject fibroblasts, resulting in strikingly abnormal mitochondrial architecture, with markedly shortened cristae. SLC25A46 also interacts with the ER membrane protein complex EMC, and phospholipid composition is altered in subject mitochondria. These results show that SLC25A46 plays a role in a mitochondrial/ER pathway that facilitates lipid transfer, and link altered mitochondrial dynamics to early‐onset neurodegenerative disease and cell fate decisions.
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spelling doaj-art-22810f2315b34bb8997e0edcfcab588d2025-08-20T03:06:00ZengSpringer NatureEMBO Molecular Medicine1757-46761757-46842016-07-01891019103810.15252/emmm.201506159SLC25A46 is required for mitochondrial lipid homeostasis and cristae maintenance and is responsible for Leigh syndromeAlexandre Janer0Julien Prudent1Vincent Paupe2Somayyeh Fahiminiya3Jacek Majewski4Nicolas Sgarioto5Christine Des Rosiers6Anik Forest7Zhen‐Yuan Lin8Anne‐Claude Gingras9Grant Mitchell10Heidi M McBride11Eric A Shoubridge12Department of Human Genetics, McGill UniversityMontreal Neurological Institute, McGill UniversityDepartment of Human Genetics, McGill UniversityDepartment of Human Genetics, McGill UniversityDepartment of Human Genetics, McGill UniversityMontreal Neurological Institute, McGill UniversityDepartment of Nutrition, Université de MontréalDepartment of Nutrition, Université de MontréalLunenfeld‐Tanenbaum Research Institute, Mount Sinai HospitalLunenfeld‐Tanenbaum Research Institute, Mount Sinai HospitalDivision of Medical Genetics, Department of Pediatrics, CHU Sainte‐Justine and Université de MontréalMontreal Neurological Institute, McGill UniversityDepartment of Human Genetics, McGill UniversityAbstract Mitochondria form a dynamic network that responds to physiological signals and metabolic stresses by altering the balance between fusion and fission. Mitochondrial fusion is orchestrated by conserved GTPases MFN1/2 and OPA1, a process coordinated in yeast by Ugo1, a mitochondrial metabolite carrier family protein. We uncovered a homozygous missense mutation in SLC25A46, the mammalian orthologue of Ugo1, in a subject with Leigh syndrome. SLC25A46 is an integral outer membrane protein that interacts with MFN2, OPA1, and the mitochondrial contact site and cristae organizing system (MICOS) complex. The subject mutation destabilizes the protein, leading to mitochondrial hyperfusion, alterations in endoplasmic reticulum (ER) morphology, impaired cellular respiration, and premature cellular senescence. The MICOS complex is disrupted in subject fibroblasts, resulting in strikingly abnormal mitochondrial architecture, with markedly shortened cristae. SLC25A46 also interacts with the ER membrane protein complex EMC, and phospholipid composition is altered in subject mitochondria. These results show that SLC25A46 plays a role in a mitochondrial/ER pathway that facilitates lipid transfer, and link altered mitochondrial dynamics to early‐onset neurodegenerative disease and cell fate decisions.https://doi.org/10.15252/emmm.201506159Leigh syndromemitochondrial architecturephospholipid transferSLC25A46
spellingShingle Alexandre Janer
Julien Prudent
Vincent Paupe
Somayyeh Fahiminiya
Jacek Majewski
Nicolas Sgarioto
Christine Des Rosiers
Anik Forest
Zhen‐Yuan Lin
Anne‐Claude Gingras
Grant Mitchell
Heidi M McBride
Eric A Shoubridge
SLC25A46 is required for mitochondrial lipid homeostasis and cristae maintenance and is responsible for Leigh syndrome
EMBO Molecular Medicine
Leigh syndrome
mitochondrial architecture
phospholipid transfer
SLC25A46
title SLC25A46 is required for mitochondrial lipid homeostasis and cristae maintenance and is responsible for Leigh syndrome
title_full SLC25A46 is required for mitochondrial lipid homeostasis and cristae maintenance and is responsible for Leigh syndrome
title_fullStr SLC25A46 is required for mitochondrial lipid homeostasis and cristae maintenance and is responsible for Leigh syndrome
title_full_unstemmed SLC25A46 is required for mitochondrial lipid homeostasis and cristae maintenance and is responsible for Leigh syndrome
title_short SLC25A46 is required for mitochondrial lipid homeostasis and cristae maintenance and is responsible for Leigh syndrome
title_sort slc25a46 is required for mitochondrial lipid homeostasis and cristae maintenance and is responsible for leigh syndrome
topic Leigh syndrome
mitochondrial architecture
phospholipid transfer
SLC25A46
url https://doi.org/10.15252/emmm.201506159
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