How do arbuscular mycorrhizal fungi enhance drought resistance of Leymus chinensis?

Abstract Background Leymus chinensis is a vital, dominant grass species in Eurasian temperate grasslands, including the Inner Mongolian steppe. L. chinensis exhibits enhanced drought tolerance through symbiosis with arbuscular mycorrhizal fungi (AMF). The physiological mechanisms behind this drought...

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Main Authors: Xiaojiang Yang, Zhen Wang, Jing Li, Paul C. Struik, Shenyi Jiang, Ke Jin, Huaibin Mu
Format: Article
Language:English
Published: BMC 2025-04-01
Series:BMC Plant Biology
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Online Access:https://doi.org/10.1186/s12870-025-06412-1
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author Xiaojiang Yang
Zhen Wang
Jing Li
Paul C. Struik
Shenyi Jiang
Ke Jin
Huaibin Mu
author_facet Xiaojiang Yang
Zhen Wang
Jing Li
Paul C. Struik
Shenyi Jiang
Ke Jin
Huaibin Mu
author_sort Xiaojiang Yang
collection DOAJ
description Abstract Background Leymus chinensis is a vital, dominant grass species in Eurasian temperate grasslands, including the Inner Mongolian steppe. L. chinensis exhibits enhanced drought tolerance through symbiosis with arbuscular mycorrhizal fungi (AMF). The physiological mechanisms behind this drought resistance need to be unraveled. A pot experiment was conducted with four inoculation treatments (inoculation with Funneliformis mosseae, with Claroideoglomus etunicatum, or with both, and no inoculation) and three drought treatments (no drought (75.00% field capacity), mild drought (56.25% field capacity), severe drought (37.50% field capacity)) to analyze how AMF enhance drought resistance of L. chinensis. Results The results showed that drought stress inhibited the growth of L. chinensis, depending on its intensity, whereas AMF inoculation significantly improved growth and alleviated the effects of drought stress. Regardless of drought conditions, AMF inoculation significantly enhanced key biochemistry parameters, including soluble sugar concentration and antioxidant enzyme activities, ultimately promoting plant productivity. Structural equation models (SEMs) further showed that the increase in biomass of L. chinensis inoculated with AMF during mild drought was primarily due to reduced catalase activity and increased cytokinin concentration by increased soluble sugar concentration. However, under severe drought, the increase in biomass of L. chinensis inoculated with AMF was associated with increased soluble sugar concentration caused by increased peroxidase activity and reduced cytokinin concentration. Conclusions The mechanisms by which AMF enhance the drought resistance of L. chinensis vary depending on the severity of drought. AMF increase the soluble sugar concentration by enhancing photosynthetic activity to improve drought resistance under mild drought. Under severe drought conditions, AMF enhance the concentration of soluble sugars in L. chinensis by further activating the expression of antioxidant enzyme genes, thereby improving its drought resistance. Additionally, C. etunicatum maintains high ectomycelium by requiring less carbon sources to efficiently absorb the residual soil moisture under severe drought, thus superiorly enhancing the drought resistance of L. chinensis. This study provides a theoretical foundation for the application of AMF fertilizer to improve the productivity of L. chinensis in arid grasslands.
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spelling doaj-art-1af97dbc456a43738832ea66a968d0db2025-08-20T02:11:41ZengBMCBMC Plant Biology1471-22292025-04-0125111410.1186/s12870-025-06412-1How do arbuscular mycorrhizal fungi enhance drought resistance of Leymus chinensis?Xiaojiang Yang0Zhen Wang1Jing Li2Paul C. Struik3Shenyi Jiang4Ke Jin5Huaibin Mu6Institute of Grassland Research, Chinese Academy of Agricultural SciencesInstitute of Grassland Research, Chinese Academy of Agricultural SciencesInstitute of Grassland Research, Chinese Academy of Agricultural SciencesCentre for Crop Systems Analysis, Department of Plant Sciences, Wageningen University and ResearchInstitute of Grassland Research, Chinese Academy of Agricultural SciencesInstitute of Grassland Research, Chinese Academy of Agricultural SciencesInstitute of Grassland Research, Chinese Academy of Agricultural SciencesAbstract Background Leymus chinensis is a vital, dominant grass species in Eurasian temperate grasslands, including the Inner Mongolian steppe. L. chinensis exhibits enhanced drought tolerance through symbiosis with arbuscular mycorrhizal fungi (AMF). The physiological mechanisms behind this drought resistance need to be unraveled. A pot experiment was conducted with four inoculation treatments (inoculation with Funneliformis mosseae, with Claroideoglomus etunicatum, or with both, and no inoculation) and three drought treatments (no drought (75.00% field capacity), mild drought (56.25% field capacity), severe drought (37.50% field capacity)) to analyze how AMF enhance drought resistance of L. chinensis. Results The results showed that drought stress inhibited the growth of L. chinensis, depending on its intensity, whereas AMF inoculation significantly improved growth and alleviated the effects of drought stress. Regardless of drought conditions, AMF inoculation significantly enhanced key biochemistry parameters, including soluble sugar concentration and antioxidant enzyme activities, ultimately promoting plant productivity. Structural equation models (SEMs) further showed that the increase in biomass of L. chinensis inoculated with AMF during mild drought was primarily due to reduced catalase activity and increased cytokinin concentration by increased soluble sugar concentration. However, under severe drought, the increase in biomass of L. chinensis inoculated with AMF was associated with increased soluble sugar concentration caused by increased peroxidase activity and reduced cytokinin concentration. Conclusions The mechanisms by which AMF enhance the drought resistance of L. chinensis vary depending on the severity of drought. AMF increase the soluble sugar concentration by enhancing photosynthetic activity to improve drought resistance under mild drought. Under severe drought conditions, AMF enhance the concentration of soluble sugars in L. chinensis by further activating the expression of antioxidant enzyme genes, thereby improving its drought resistance. Additionally, C. etunicatum maintains high ectomycelium by requiring less carbon sources to efficiently absorb the residual soil moisture under severe drought, thus superiorly enhancing the drought resistance of L. chinensis. This study provides a theoretical foundation for the application of AMF fertilizer to improve the productivity of L. chinensis in arid grasslands.https://doi.org/10.1186/s12870-025-06412-1Arbuscular mycorrhizal fungiPlant physiology and biochemistryLeymus chinensisDrought resistance mechanism
spellingShingle Xiaojiang Yang
Zhen Wang
Jing Li
Paul C. Struik
Shenyi Jiang
Ke Jin
Huaibin Mu
How do arbuscular mycorrhizal fungi enhance drought resistance of Leymus chinensis?
BMC Plant Biology
Arbuscular mycorrhizal fungi
Plant physiology and biochemistry
Leymus chinensis
Drought resistance mechanism
title How do arbuscular mycorrhizal fungi enhance drought resistance of Leymus chinensis?
title_full How do arbuscular mycorrhizal fungi enhance drought resistance of Leymus chinensis?
title_fullStr How do arbuscular mycorrhizal fungi enhance drought resistance of Leymus chinensis?
title_full_unstemmed How do arbuscular mycorrhizal fungi enhance drought resistance of Leymus chinensis?
title_short How do arbuscular mycorrhizal fungi enhance drought resistance of Leymus chinensis?
title_sort how do arbuscular mycorrhizal fungi enhance drought resistance of leymus chinensis
topic Arbuscular mycorrhizal fungi
Plant physiology and biochemistry
Leymus chinensis
Drought resistance mechanism
url https://doi.org/10.1186/s12870-025-06412-1
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