BrnQ Branched-Chain Amino Acid Transporters Influence Toxin Production by, but Not Growth of, <i>Clostridium perfringens</i> Type A Strain ATCC3624

By producing alpha toxin (PLC) and perfringolysin O (PFO), <i>Clostridium perfringens</i> type A strains are the most common cause of traumatic gas gangrene. <i>C. perfringens</i> cannot synthesize branched-chain amino acids (BCAAs), so BCAA transporters are essential for <...

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Main Authors: Jihong Li, Iman Mehdizadeh Gohari, Isabella Zhang, Bruce A. McClane
Format: Article
Language:English
Published: MDPI AG 2025-04-01
Series:Toxins
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Online Access:https://www.mdpi.com/2072-6651/17/4/187
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author Jihong Li
Iman Mehdizadeh Gohari
Isabella Zhang
Bruce A. McClane
author_facet Jihong Li
Iman Mehdizadeh Gohari
Isabella Zhang
Bruce A. McClane
author_sort Jihong Li
collection DOAJ
description By producing alpha toxin (PLC) and perfringolysin O (PFO), <i>Clostridium perfringens</i> type A strains are the most common cause of traumatic gas gangrene. <i>C. perfringens</i> cannot synthesize branched-chain amino acids (BCAAs), so BCAA transporters are essential for <i>C. perfringens</i> growth and survival. <i>C. perfringens</i> type A strain ATCC3624 encodes the BrnQ1, BrnQ2, and BrnQ3 BCAA transporters. RT-PCR analyses showed that, with increasing culture time in TY broth, <i>brnQ2</i> and <i>brnQ3</i> expression levels remained stable but <i>brnQ1</i> expression levels declined. Single null mutants unable to produce one of the BrnQ proteins grew and survived similarly as wild type. However, these mutants all showed altered PLC production, especially in the early culture stage, and those effects were reversible by complementation. Therefore, the presence of BrnQ proteins impacts toxin production levels, even though they are not necessary for growth. Interestingly, a triple mutant that was unable to produce any BrnQ protein also grew similarly as ATCC3624. Since BCAA uptake is essential for <i>C. perfringens</i>, this strain must produce another (still to be identified) BCAA transporter.
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spelling doaj-art-19c02e4c557948bb87e87cc5fbd2d06a2025-08-20T02:18:10ZengMDPI AGToxins2072-66512025-04-0117418710.3390/toxins17040187BrnQ Branched-Chain Amino Acid Transporters Influence Toxin Production by, but Not Growth of, <i>Clostridium perfringens</i> Type A Strain ATCC3624Jihong Li0Iman Mehdizadeh Gohari1Isabella Zhang2Bruce A. McClane3Department of Microbiology and Molecular Genetics, School of Medicine, University of Pittsburgh, Pittsburgh, PA 15219, USADepartment of Microbiology and Molecular Genetics, School of Medicine, University of Pittsburgh, Pittsburgh, PA 15219, USADepartment of Microbiology and Molecular Genetics, School of Medicine, University of Pittsburgh, Pittsburgh, PA 15219, USADepartment of Microbiology and Molecular Genetics, School of Medicine, University of Pittsburgh, Pittsburgh, PA 15219, USABy producing alpha toxin (PLC) and perfringolysin O (PFO), <i>Clostridium perfringens</i> type A strains are the most common cause of traumatic gas gangrene. <i>C. perfringens</i> cannot synthesize branched-chain amino acids (BCAAs), so BCAA transporters are essential for <i>C. perfringens</i> growth and survival. <i>C. perfringens</i> type A strain ATCC3624 encodes the BrnQ1, BrnQ2, and BrnQ3 BCAA transporters. RT-PCR analyses showed that, with increasing culture time in TY broth, <i>brnQ2</i> and <i>brnQ3</i> expression levels remained stable but <i>brnQ1</i> expression levels declined. Single null mutants unable to produce one of the BrnQ proteins grew and survived similarly as wild type. However, these mutants all showed altered PLC production, especially in the early culture stage, and those effects were reversible by complementation. Therefore, the presence of BrnQ proteins impacts toxin production levels, even though they are not necessary for growth. Interestingly, a triple mutant that was unable to produce any BrnQ protein also grew similarly as ATCC3624. Since BCAA uptake is essential for <i>C. perfringens</i>, this strain must produce another (still to be identified) BCAA transporter.https://www.mdpi.com/2072-6651/17/4/187<i>Clostridium perfringens</i>BrnQ transportersalpha toxinperfringolysin Oregulation
spellingShingle Jihong Li
Iman Mehdizadeh Gohari
Isabella Zhang
Bruce A. McClane
BrnQ Branched-Chain Amino Acid Transporters Influence Toxin Production by, but Not Growth of, <i>Clostridium perfringens</i> Type A Strain ATCC3624
Toxins
<i>Clostridium perfringens</i>
BrnQ transporters
alpha toxin
perfringolysin O
regulation
title BrnQ Branched-Chain Amino Acid Transporters Influence Toxin Production by, but Not Growth of, <i>Clostridium perfringens</i> Type A Strain ATCC3624
title_full BrnQ Branched-Chain Amino Acid Transporters Influence Toxin Production by, but Not Growth of, <i>Clostridium perfringens</i> Type A Strain ATCC3624
title_fullStr BrnQ Branched-Chain Amino Acid Transporters Influence Toxin Production by, but Not Growth of, <i>Clostridium perfringens</i> Type A Strain ATCC3624
title_full_unstemmed BrnQ Branched-Chain Amino Acid Transporters Influence Toxin Production by, but Not Growth of, <i>Clostridium perfringens</i> Type A Strain ATCC3624
title_short BrnQ Branched-Chain Amino Acid Transporters Influence Toxin Production by, but Not Growth of, <i>Clostridium perfringens</i> Type A Strain ATCC3624
title_sort brnq branched chain amino acid transporters influence toxin production by but not growth of i clostridium perfringens i type a strain atcc3624
topic <i>Clostridium perfringens</i>
BrnQ transporters
alpha toxin
perfringolysin O
regulation
url https://www.mdpi.com/2072-6651/17/4/187
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