TDP-43 overexpression in the hypothalamus drives neuropathology, dysregulates metabolism and impairs behavior in mice

Abstract TAR DNA-binding protein 43 (TDP-43) pathology is linked to the neurodegenerative disorders amyotrophic lateral sclerosis (ALS), frontotemporal dementia (FTD) and Huntington disease (HD). Dysregulation of metabolism and emotion is shared across these disorders and may be caused by hypothalam...

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Main Authors: Sofia Bergh, Nicolas Casadei, Sanaz Gabery, Oskar Simonsson, João M. N. Duarte, Deniz Kirik, Huu Phuc Nguyen, Åsa Petersén
Format: Article
Language:English
Published: BMC 2025-05-01
Series:Acta Neuropathologica Communications
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Online Access:https://doi.org/10.1186/s40478-025-02018-8
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author Sofia Bergh
Nicolas Casadei
Sanaz Gabery
Oskar Simonsson
João M. N. Duarte
Deniz Kirik
Huu Phuc Nguyen
Åsa Petersén
author_facet Sofia Bergh
Nicolas Casadei
Sanaz Gabery
Oskar Simonsson
João M. N. Duarte
Deniz Kirik
Huu Phuc Nguyen
Åsa Petersén
author_sort Sofia Bergh
collection DOAJ
description Abstract TAR DNA-binding protein 43 (TDP-43) pathology is linked to the neurodegenerative disorders amyotrophic lateral sclerosis (ALS), frontotemporal dementia (FTD) and Huntington disease (HD). Dysregulation of metabolism and emotion is shared across these disorders and may be caused by hypothalamic pathology. Inclusions with TDP-43 are present in the hypothalamus in clinical ALS, as well as selective loss of hypothalamic neurons expressing the metabolism and emotion regulating neuropeptides hypocretin (orexin), melanin-concentrating hormone (MCH) and oxytocin. We aimed to investigate whether there is a casual link between the effects of TDP-43 in the hypothalamus and the development of neuropathology, as well as changes in metabolism and behavior. We generated an adeno-associated viral (AAV) vector expressing human TDP-43 under the neuronal-specific synapsin promoter, which was injected bilaterally into the hypothalamus of wild-type FVB/N mice. TDP-43 overexpression resulted in hypothalamic pathology in a dose-dependent fashion replicating clinical pathology with hypothalamic atrophy and loss of hypocretin-, MCH- and oxytocin-expressing neurons. Nuclear and cytoplasmic inclusions of TDP-43 were found in the hypothalamus. Mice overexpressing TDP-43 in the hypothalamus developed metabolic dysregulation with hyperglycaemia independent of food intake. Additionally, mice overexpressing TDP-43 in the hypothalamus exhibited reduced motor activity and nesting ability, suggesting the development of an apathy-like phenotype. Taken together, AAV-vector mediated TDP-43 overexpression in the hypothalamus leads to neuropathology with the development of metabolic dysfunction and apathy-like behavior. These results indicate that TDP-43 can exert direct pathological effects in the hypothalamus, which may contribute to the development of the non-motor phenotype in TDP-43 proteinopathies.
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spelling doaj-art-15c02dad965c4aefabe3818e4be24e6b2025-08-20T02:00:13ZengBMCActa Neuropathologica Communications2051-59602025-05-0113112110.1186/s40478-025-02018-8TDP-43 overexpression in the hypothalamus drives neuropathology, dysregulates metabolism and impairs behavior in miceSofia Bergh0Nicolas Casadei1Sanaz Gabery2Oskar Simonsson3João M. N. Duarte4Deniz Kirik5Huu Phuc Nguyen6Åsa Petersén7Translational Neuroendocrine Research Unit (TNU), Department of Experimental Medical Science, Lund UniversityInstitute of Medical Genetics and Applied Genomics, University of TübingenTranslational Neuroendocrine Research Unit (TNU), Department of Experimental Medical Science, Lund UniversityTranslational Neuroendocrine Research Unit (TNU), Department of Experimental Medical Science, Lund UniversityDiabetes and Brain Function Unit, Department of Experimental Medical Science, Lund UniversityBrain Repair and Imaging in Neural Systems (BRAINS), Department of Experimental Medical Science, Lund UniversityDepartment of Human Genetics, Medical Faculty, Ruhr University BochumTranslational Neuroendocrine Research Unit (TNU), Department of Experimental Medical Science, Lund UniversityAbstract TAR DNA-binding protein 43 (TDP-43) pathology is linked to the neurodegenerative disorders amyotrophic lateral sclerosis (ALS), frontotemporal dementia (FTD) and Huntington disease (HD). Dysregulation of metabolism and emotion is shared across these disorders and may be caused by hypothalamic pathology. Inclusions with TDP-43 are present in the hypothalamus in clinical ALS, as well as selective loss of hypothalamic neurons expressing the metabolism and emotion regulating neuropeptides hypocretin (orexin), melanin-concentrating hormone (MCH) and oxytocin. We aimed to investigate whether there is a casual link between the effects of TDP-43 in the hypothalamus and the development of neuropathology, as well as changes in metabolism and behavior. We generated an adeno-associated viral (AAV) vector expressing human TDP-43 under the neuronal-specific synapsin promoter, which was injected bilaterally into the hypothalamus of wild-type FVB/N mice. TDP-43 overexpression resulted in hypothalamic pathology in a dose-dependent fashion replicating clinical pathology with hypothalamic atrophy and loss of hypocretin-, MCH- and oxytocin-expressing neurons. Nuclear and cytoplasmic inclusions of TDP-43 were found in the hypothalamus. Mice overexpressing TDP-43 in the hypothalamus developed metabolic dysregulation with hyperglycaemia independent of food intake. Additionally, mice overexpressing TDP-43 in the hypothalamus exhibited reduced motor activity and nesting ability, suggesting the development of an apathy-like phenotype. Taken together, AAV-vector mediated TDP-43 overexpression in the hypothalamus leads to neuropathology with the development of metabolic dysfunction and apathy-like behavior. These results indicate that TDP-43 can exert direct pathological effects in the hypothalamus, which may contribute to the development of the non-motor phenotype in TDP-43 proteinopathies.https://doi.org/10.1186/s40478-025-02018-8(4 to 6): TDP-43HypothalamusMetabolismAmyotrophic lateral sclerosisFrontotemporal dementia
spellingShingle Sofia Bergh
Nicolas Casadei
Sanaz Gabery
Oskar Simonsson
João M. N. Duarte
Deniz Kirik
Huu Phuc Nguyen
Åsa Petersén
TDP-43 overexpression in the hypothalamus drives neuropathology, dysregulates metabolism and impairs behavior in mice
Acta Neuropathologica Communications
(4 to 6): TDP-43
Hypothalamus
Metabolism
Amyotrophic lateral sclerosis
Frontotemporal dementia
title TDP-43 overexpression in the hypothalamus drives neuropathology, dysregulates metabolism and impairs behavior in mice
title_full TDP-43 overexpression in the hypothalamus drives neuropathology, dysregulates metabolism and impairs behavior in mice
title_fullStr TDP-43 overexpression in the hypothalamus drives neuropathology, dysregulates metabolism and impairs behavior in mice
title_full_unstemmed TDP-43 overexpression in the hypothalamus drives neuropathology, dysregulates metabolism and impairs behavior in mice
title_short TDP-43 overexpression in the hypothalamus drives neuropathology, dysregulates metabolism and impairs behavior in mice
title_sort tdp 43 overexpression in the hypothalamus drives neuropathology dysregulates metabolism and impairs behavior in mice
topic (4 to 6): TDP-43
Hypothalamus
Metabolism
Amyotrophic lateral sclerosis
Frontotemporal dementia
url https://doi.org/10.1186/s40478-025-02018-8
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