Humerus shape evolved in cetaceans under relaxed selection and random drift

Abstract Mammalian humerus shape is mostly predicted by loading. This constraint is lifted for cetaceans and, therefore, can reflect other, specific evolutionary trends. To test this hypothesis, the three-dimensional shape of the humerus for 36 taxa of extinct and extant cetaceans was analyzed. The...

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Main Authors: Maria Ghazali, Svitozar Davydenko, Valeriia Telizhenko, Pavlo Otriazhyi, Karina Vishnyakova, Maia Bukhsianidze, Azucena Solis-Añorve, Igor Dzeverin, Pavel Gol’din
Format: Article
Language:English
Published: Nature Portfolio 2025-03-01
Series:Communications Biology
Online Access:https://doi.org/10.1038/s42003-025-07952-w
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author Maria Ghazali
Svitozar Davydenko
Valeriia Telizhenko
Pavlo Otriazhyi
Karina Vishnyakova
Maia Bukhsianidze
Azucena Solis-Añorve
Igor Dzeverin
Pavel Gol’din
author_facet Maria Ghazali
Svitozar Davydenko
Valeriia Telizhenko
Pavlo Otriazhyi
Karina Vishnyakova
Maia Bukhsianidze
Azucena Solis-Añorve
Igor Dzeverin
Pavel Gol’din
author_sort Maria Ghazali
collection DOAJ
description Abstract Mammalian humerus shape is mostly predicted by loading. This constraint is lifted for cetaceans and, therefore, can reflect other, specific evolutionary trends. To test this hypothesis, the three-dimensional shape of the humerus for 36 taxa of extinct and extant cetaceans was analyzed. The shape variance concurs with the evolution of diverging baleen and toothed whales and is independent of allometry. It shows traits associated with aquatic locomotion: humeral head twist, diaphysis shortening and straightening, and epiphyses enlargement. Also, changes in the anatomy of the humeral head and greater tubercle, modularity and integration patterns are associated with phylogeny. We suggest relaxed negative selection and random drift as the likely main evolutionary patterns for the evolution of the humerus shape. The evolution of genes regulating the humerus shape corresponds to the anatomical transformations and shows relaxed selection, site-level positive selection and nonsense mutations in cetaceans. Overall, the observed pattern well illustrates a “fly in a tube model” and shows its importance as a framework of emerging evolutionary innovations.
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spelling doaj-art-13dec9ecef6b46318ca95a56ee3901b22025-08-20T02:10:14ZengNature PortfolioCommunications Biology2399-36422025-03-018111510.1038/s42003-025-07952-wHumerus shape evolved in cetaceans under relaxed selection and random driftMaria Ghazali0Svitozar Davydenko1Valeriia Telizhenko2Pavlo Otriazhyi3Karina Vishnyakova4Maia Bukhsianidze5Azucena Solis-Añorve6Igor Dzeverin7Pavel Gol’din8Schmalhausen Institute of Zoology, National Academy of Sciences of UkraineSchmalhausen Institute of Zoology, National Academy of Sciences of UkraineSchmalhausen Institute of Zoology, National Academy of Sciences of UkraineSchmalhausen Institute of Zoology, National Academy of Sciences of UkraineUkrainian Scientific Centre of Ecology of the SeaGeorgian National MuseumAutonomous University of Baja California SurSchmalhausen Institute of Zoology, National Academy of Sciences of UkraineSchmalhausen Institute of Zoology, National Academy of Sciences of UkraineAbstract Mammalian humerus shape is mostly predicted by loading. This constraint is lifted for cetaceans and, therefore, can reflect other, specific evolutionary trends. To test this hypothesis, the three-dimensional shape of the humerus for 36 taxa of extinct and extant cetaceans was analyzed. The shape variance concurs with the evolution of diverging baleen and toothed whales and is independent of allometry. It shows traits associated with aquatic locomotion: humeral head twist, diaphysis shortening and straightening, and epiphyses enlargement. Also, changes in the anatomy of the humeral head and greater tubercle, modularity and integration patterns are associated with phylogeny. We suggest relaxed negative selection and random drift as the likely main evolutionary patterns for the evolution of the humerus shape. The evolution of genes regulating the humerus shape corresponds to the anatomical transformations and shows relaxed selection, site-level positive selection and nonsense mutations in cetaceans. Overall, the observed pattern well illustrates a “fly in a tube model” and shows its importance as a framework of emerging evolutionary innovations.https://doi.org/10.1038/s42003-025-07952-w
spellingShingle Maria Ghazali
Svitozar Davydenko
Valeriia Telizhenko
Pavlo Otriazhyi
Karina Vishnyakova
Maia Bukhsianidze
Azucena Solis-Añorve
Igor Dzeverin
Pavel Gol’din
Humerus shape evolved in cetaceans under relaxed selection and random drift
Communications Biology
title Humerus shape evolved in cetaceans under relaxed selection and random drift
title_full Humerus shape evolved in cetaceans under relaxed selection and random drift
title_fullStr Humerus shape evolved in cetaceans under relaxed selection and random drift
title_full_unstemmed Humerus shape evolved in cetaceans under relaxed selection and random drift
title_short Humerus shape evolved in cetaceans under relaxed selection and random drift
title_sort humerus shape evolved in cetaceans under relaxed selection and random drift
url https://doi.org/10.1038/s42003-025-07952-w
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