The Venom Proteome of the Ecologically Divergent Australian Elapid, Southern Death Adder <i>Acanthophis antarcticus</i>

The composition of Australian snake venoms is the least well-known of any continent. We characterised the venom proteome of the southern death adder <i>Acanthophis antarcticus</i>—one of the world’s most morphologically and ecologically divergent elapids. Using a combined bottom-up prote...

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Main Authors: Theo Tasoulis, C. Ruth Wang, Shaun Ellis, Tara L. Pukala, Joanna Sumner, Kate Murphy, Nathan Dunstan, Geoffrey K. Isbister
Format: Article
Language:English
Published: MDPI AG 2025-07-01
Series:Toxins
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Online Access:https://www.mdpi.com/2072-6651/17/7/352
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author Theo Tasoulis
C. Ruth Wang
Shaun Ellis
Tara L. Pukala
Joanna Sumner
Kate Murphy
Nathan Dunstan
Geoffrey K. Isbister
author_facet Theo Tasoulis
C. Ruth Wang
Shaun Ellis
Tara L. Pukala
Joanna Sumner
Kate Murphy
Nathan Dunstan
Geoffrey K. Isbister
author_sort Theo Tasoulis
collection DOAJ
description The composition of Australian snake venoms is the least well-known of any continent. We characterised the venom proteome of the southern death adder <i>Acanthophis antarcticus</i>—one of the world’s most morphologically and ecologically divergent elapids. Using a combined bottom-up proteomic and venom gland transcriptomic approach employing reverse-phase chromatographic and gel electrophoretic fractionation strategies in the bottom-up proteomic workflow, we characterised 92.8% of the venom, comprising twelve different toxin identification hits belonging to seven toxin families. The most abundant protein family was three-finger toxins (3FTxs; 59.8% whole venom), consisting mostly of one long-chain neurotoxin, alpha-elapitoxin-Aa2b making up 59% of the venom and two proteoforms of another long-chain neurotoxin. Phospholipase A<sub>2</sub>s (PLA<sub>2</sub>s) were the second most abundant, with four different toxins making up 22.5% of the venom. One toxin was similar to two previous non-neurotoxic PLA<sub>2</sub>s, making up 16% of the venom. The remaining protein families present were CTL (3.6%), NGF (2.5%), CRiSP (1.8%), LAAO (1.4%), and AChE (0.8%). <i>A. antarcticus</i> is the first Australian elapid characterised that has a 3FTx dominant venom, a composition typical of elapids on other continents, particularly cobras <i>Naja</i> sp. The fact that <i>A. antarcticus</i> has a venom composition similar to cobra venom while having a viper-like ecology illustrates that similar venom expressions can evolve independently of ecology. The predominance of post-synaptic neurotoxins (3FTxs) and pre-synaptic neurotoxins (PLA<sub>2</sub>) is consistent with the neurotoxic clinical effects of envenomation in humans.
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spelling doaj-art-106bb56f569947e7bede1de1d289f82a2025-08-20T02:47:18ZengMDPI AGToxins2072-66512025-07-0117735210.3390/toxins17070352The Venom Proteome of the Ecologically Divergent Australian Elapid, Southern Death Adder <i>Acanthophis antarcticus</i>Theo Tasoulis0C. Ruth Wang1Shaun Ellis2Tara L. Pukala3Joanna Sumner4Kate Murphy5Nathan Dunstan6Geoffrey K. Isbister7Clinical Toxicology Research Group, University of Newcastle, Newcastle, NSW 2308, AustraliaDepartment of Chemistry, University of Adelaide, Adelaide, SA 5000, AustraliaDepartment of Chemistry, University of Adelaide, Adelaide, SA 5000, AustraliaDepartment of Chemistry, University of Adelaide, Adelaide, SA 5000, AustraliaMuseums Victoria, Melbourne, VIC 5053, AustraliaClinical Toxicology Research Group, University of Newcastle, Newcastle, NSW 2308, AustraliaVenom Supplies, Tanunda, Adelaide, SA 5352, AustraliaClinical Toxicology Research Group, University of Newcastle, Newcastle, NSW 2308, AustraliaThe composition of Australian snake venoms is the least well-known of any continent. We characterised the venom proteome of the southern death adder <i>Acanthophis antarcticus</i>—one of the world’s most morphologically and ecologically divergent elapids. Using a combined bottom-up proteomic and venom gland transcriptomic approach employing reverse-phase chromatographic and gel electrophoretic fractionation strategies in the bottom-up proteomic workflow, we characterised 92.8% of the venom, comprising twelve different toxin identification hits belonging to seven toxin families. The most abundant protein family was three-finger toxins (3FTxs; 59.8% whole venom), consisting mostly of one long-chain neurotoxin, alpha-elapitoxin-Aa2b making up 59% of the venom and two proteoforms of another long-chain neurotoxin. Phospholipase A<sub>2</sub>s (PLA<sub>2</sub>s) were the second most abundant, with four different toxins making up 22.5% of the venom. One toxin was similar to two previous non-neurotoxic PLA<sub>2</sub>s, making up 16% of the venom. The remaining protein families present were CTL (3.6%), NGF (2.5%), CRiSP (1.8%), LAAO (1.4%), and AChE (0.8%). <i>A. antarcticus</i> is the first Australian elapid characterised that has a 3FTx dominant venom, a composition typical of elapids on other continents, particularly cobras <i>Naja</i> sp. The fact that <i>A. antarcticus</i> has a venom composition similar to cobra venom while having a viper-like ecology illustrates that similar venom expressions can evolve independently of ecology. The predominance of post-synaptic neurotoxins (3FTxs) and pre-synaptic neurotoxins (PLA<sub>2</sub>) is consistent with the neurotoxic clinical effects of envenomation in humans.https://www.mdpi.com/2072-6651/17/7/352snake venomtoxinsproteomicsvenomephospholipaseecology
spellingShingle Theo Tasoulis
C. Ruth Wang
Shaun Ellis
Tara L. Pukala
Joanna Sumner
Kate Murphy
Nathan Dunstan
Geoffrey K. Isbister
The Venom Proteome of the Ecologically Divergent Australian Elapid, Southern Death Adder <i>Acanthophis antarcticus</i>
Toxins
snake venom
toxins
proteomics
venome
phospholipase
ecology
title The Venom Proteome of the Ecologically Divergent Australian Elapid, Southern Death Adder <i>Acanthophis antarcticus</i>
title_full The Venom Proteome of the Ecologically Divergent Australian Elapid, Southern Death Adder <i>Acanthophis antarcticus</i>
title_fullStr The Venom Proteome of the Ecologically Divergent Australian Elapid, Southern Death Adder <i>Acanthophis antarcticus</i>
title_full_unstemmed The Venom Proteome of the Ecologically Divergent Australian Elapid, Southern Death Adder <i>Acanthophis antarcticus</i>
title_short The Venom Proteome of the Ecologically Divergent Australian Elapid, Southern Death Adder <i>Acanthophis antarcticus</i>
title_sort venom proteome of the ecologically divergent australian elapid southern death adder i acanthophis antarcticus i
topic snake venom
toxins
proteomics
venome
phospholipase
ecology
url https://www.mdpi.com/2072-6651/17/7/352
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