Deletion of RAP1 affects iron homeostasis, azole resistance, and virulence in Candida albicans

ABSTRACT Rap1 is a DNA-binding protein conserved from yeast to mammals for its role in telomeric maintenance. Here, to explore additional functions of Candida albicans Rap1, we performed RNA sequencing analysis. Experimental validations further showed that Rap1 plays a role in iron regulation, espec...

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Main Authors: Min-Chi Yang, Wei-Luen Huang, Hsuan-Yu Chen, Shin-Huey Lin, Yu-Shan Chang, Kuo-Yun Tseng, Hsiu-Jung Lo, I-Ching Wang, Chi-Jan Lin, Chung-Yu Lan
Format: Article
Language:English
Published: American Society for Microbiology 2025-05-01
Series:mSphere
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Online Access:https://journals.asm.org/doi/10.1128/msphere.00155-25
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author Min-Chi Yang
Wei-Luen Huang
Hsuan-Yu Chen
Shin-Huey Lin
Yu-Shan Chang
Kuo-Yun Tseng
Hsiu-Jung Lo
I-Ching Wang
Chi-Jan Lin
Chung-Yu Lan
author_facet Min-Chi Yang
Wei-Luen Huang
Hsuan-Yu Chen
Shin-Huey Lin
Yu-Shan Chang
Kuo-Yun Tseng
Hsiu-Jung Lo
I-Ching Wang
Chi-Jan Lin
Chung-Yu Lan
author_sort Min-Chi Yang
collection DOAJ
description ABSTRACT Rap1 is a DNA-binding protein conserved from yeast to mammals for its role in telomeric maintenance. Here, to explore additional functions of Candida albicans Rap1, we performed RNA sequencing analysis. Experimental validations further showed that Rap1 plays a role in iron regulation, especially under low-iron conditions. Moreover, Rap1 was involved in iron acquisition and modulation of iron-related genes. Rap1 was found to be associated with fluconazole resistance in a low-iron condition. Finally, we demonstrated that the deletion of RAP1 leads to reduced C. albicans virulence in a mouse model of infection. Together, this study reveals new functions of C. albicans Rap1, particularly in iron homeostasis, azole resistance, and virulence.IMPORTANCECandida albicans is an important pathogenic fungus that can cause superficial to life-threatening infections. Iron is essential for almost all organisms, yet it is highly restricted within the human host to defend against pathogens. To grow and survive in the iron-limited host environment, C. albicans has evolved multiple iron acquisition mechanisms. Understanding the regulation of iron homeostasis is, therefore, critical for elucidating C. albicans pathogenesis and virulence. This study explores the novel functions of C. albicans Rap1, with a focus on its contribution to iron acquisition and utilization. Our findings further highlight how iron availability impacts antifungal resistance and virulence through Rap1, providing insight into the complex iron regulatory machinery of C. albicans.
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spelling doaj-art-0d438870e23c48aa8cb2da80c03f5b6b2025-08-20T01:56:25ZengAmerican Society for MicrobiologymSphere2379-50422025-05-0110510.1128/msphere.00155-25Deletion of RAP1 affects iron homeostasis, azole resistance, and virulence in Candida albicansMin-Chi Yang0Wei-Luen Huang1Hsuan-Yu Chen2Shin-Huey Lin3Yu-Shan Chang4Kuo-Yun Tseng5Hsiu-Jung Lo6I-Ching Wang7Chi-Jan Lin8Chung-Yu Lan9Institute of Molecular and Cellular Biology, National Tsing Hua University, Hsinchu, TaiwanInstitute of Molecular and Cellular Biology, National Tsing Hua University, Hsinchu, TaiwanInstitute of Molecular and Cellular Biology, National Tsing Hua University, Hsinchu, TaiwanInstitute of Biotechnology, National Tsing Hua University, Hsinchu, TaiwanInstitute of Molecular Biology, National Chung Hsing University, Taichung, TaiwanInstitute of Molecular and Cellular Biology, National Tsing Hua University, Hsinchu, TaiwanTaiwan Mycology Reference Center, National Institute of Infectious Diseases and Vaccinology, National Health Research Institutes, Miaoli County, TaiwanInstitute of Biotechnology, National Tsing Hua University, Hsinchu, TaiwanInstitute of Molecular Biology, National Chung Hsing University, Taichung, TaiwanInstitute of Molecular and Cellular Biology, National Tsing Hua University, Hsinchu, TaiwanABSTRACT Rap1 is a DNA-binding protein conserved from yeast to mammals for its role in telomeric maintenance. Here, to explore additional functions of Candida albicans Rap1, we performed RNA sequencing analysis. Experimental validations further showed that Rap1 plays a role in iron regulation, especially under low-iron conditions. Moreover, Rap1 was involved in iron acquisition and modulation of iron-related genes. Rap1 was found to be associated with fluconazole resistance in a low-iron condition. Finally, we demonstrated that the deletion of RAP1 leads to reduced C. albicans virulence in a mouse model of infection. Together, this study reveals new functions of C. albicans Rap1, particularly in iron homeostasis, azole resistance, and virulence.IMPORTANCECandida albicans is an important pathogenic fungus that can cause superficial to life-threatening infections. Iron is essential for almost all organisms, yet it is highly restricted within the human host to defend against pathogens. To grow and survive in the iron-limited host environment, C. albicans has evolved multiple iron acquisition mechanisms. Understanding the regulation of iron homeostasis is, therefore, critical for elucidating C. albicans pathogenesis and virulence. This study explores the novel functions of C. albicans Rap1, with a focus on its contribution to iron acquisition and utilization. Our findings further highlight how iron availability impacts antifungal resistance and virulence through Rap1, providing insight into the complex iron regulatory machinery of C. albicans.https://journals.asm.org/doi/10.1128/msphere.00155-25Candida albicansRap1iron acquisitioniron utilizationvirulencefluconazole susceptibility
spellingShingle Min-Chi Yang
Wei-Luen Huang
Hsuan-Yu Chen
Shin-Huey Lin
Yu-Shan Chang
Kuo-Yun Tseng
Hsiu-Jung Lo
I-Ching Wang
Chi-Jan Lin
Chung-Yu Lan
Deletion of RAP1 affects iron homeostasis, azole resistance, and virulence in Candida albicans
mSphere
Candida albicans
Rap1
iron acquisition
iron utilization
virulence
fluconazole susceptibility
title Deletion of RAP1 affects iron homeostasis, azole resistance, and virulence in Candida albicans
title_full Deletion of RAP1 affects iron homeostasis, azole resistance, and virulence in Candida albicans
title_fullStr Deletion of RAP1 affects iron homeostasis, azole resistance, and virulence in Candida albicans
title_full_unstemmed Deletion of RAP1 affects iron homeostasis, azole resistance, and virulence in Candida albicans
title_short Deletion of RAP1 affects iron homeostasis, azole resistance, and virulence in Candida albicans
title_sort deletion of rap1 affects iron homeostasis azole resistance and virulence in candida albicans
topic Candida albicans
Rap1
iron acquisition
iron utilization
virulence
fluconazole susceptibility
url https://journals.asm.org/doi/10.1128/msphere.00155-25
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