Genetic Deficiency of Hyaluronan Synthase 2 in the Developing Limb Mesenchyme Impairs Postnatal Synovial Joint Formation

Hyaluronan, a key component of the extracellular matrix, plays a crucial role in joint development and maintenance. In order to determine the role of hyaluronan function in joint development and homeostasis, conditional loss-of-function experiments of Hyaluronan Synthase 2 (<i>Has2</i>)...

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Main Authors: Yingcui Li, Alexander Tress, Peter Maye, Kemar Edwards, Asiona Findletar, Nathaniel A. Dyment, Yu Yamaguchi, David W. Rowe, Gengyun Le-Chan, Sunny S. K. Chan, Kevin W.-H. Lo
Format: Article
Language:English
Published: MDPI AG 2025-05-01
Series:Biomedicines
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Online Access:https://www.mdpi.com/2227-9059/13/6/1324
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author Yingcui Li
Alexander Tress
Peter Maye
Kemar Edwards
Asiona Findletar
Nathaniel A. Dyment
Yu Yamaguchi
David W. Rowe
Gengyun Le-Chan
Sunny S. K. Chan
Kevin W.-H. Lo
author_facet Yingcui Li
Alexander Tress
Peter Maye
Kemar Edwards
Asiona Findletar
Nathaniel A. Dyment
Yu Yamaguchi
David W. Rowe
Gengyun Le-Chan
Sunny S. K. Chan
Kevin W.-H. Lo
author_sort Yingcui Li
collection DOAJ
description Hyaluronan, a key component of the extracellular matrix, plays a crucial role in joint development and maintenance. In order to determine the role of hyaluronan function in joint development and homeostasis, conditional loss-of-function experiments of Hyaluronan Synthase 2 (<i>Has2</i>) were carried out in mice. <i>Has2</i> depletion in limb mesenchymal cells led to severely shortened limbs with appendicular joints that are deformed, decreased proteoglycan content as characterized by Safranin-O staining, and severely pitted epiphyseal ends of long bones and deformed joints as viewed by micro-CT reconstructions. The embryonic deletion of <i>Has2</i> in mesoderm mesenchyme of limbs by Prx1-Cre confirmed its involvement in joint development, while in situ hybridization and hyaluronan staining confirmed <i>Has2</i> expression and abundant accumulation of hyaluronan in the onset of joint formation, the joint interzone. These findings position <i>Has2</i> as the main hyaluronan-making enzyme in articular cartilage and highlight its essential function in joint formation and retention of proteoglycans of the extracellular matrix of the cartilage.
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spelling doaj-art-0acaa9f914b3470daab85cdce6455bdb2025-08-20T02:24:26ZengMDPI AGBiomedicines2227-90592025-05-01136132410.3390/biomedicines13061324Genetic Deficiency of Hyaluronan Synthase 2 in the Developing Limb Mesenchyme Impairs Postnatal Synovial Joint FormationYingcui Li0Alexander Tress1Peter Maye2Kemar Edwards3Asiona Findletar4Nathaniel A. Dyment5Yu Yamaguchi6David W. Rowe7Gengyun Le-Chan8Sunny S. K. Chan9Kevin W.-H. Lo10Department of Biology, College of Arts and Sciences, University of Hartford, 200 Bloomfield Avenue, West Hartford, CT 06117, USADepartment of Biology, College of Arts and Sciences, University of Hartford, 200 Bloomfield Avenue, West Hartford, CT 06117, USACenter for Regenerative Medicine and Skeletal Development, University of Connecticut Health Center, Farmington, CT 06032, USADepartment of Biology, College of Arts and Sciences, University of Hartford, 200 Bloomfield Avenue, West Hartford, CT 06117, USADepartment of Biology, College of Arts and Sciences, University of Hartford, 200 Bloomfield Avenue, West Hartford, CT 06117, USADepartment of Orthopedic Surgery, University of Pennsylvania, Philadelphia, PA 19104, USAHuman Genetics Program, Sanford Burnham Prebys Medical Discovery Institute, La Jolla, CA 92037, USACenter for Regenerative Medicine and Skeletal Development, University of Connecticut Health Center, Farmington, CT 06032, USADepartment of Biology, College of Arts and Sciences, University of Hartford, 200 Bloomfield Avenue, West Hartford, CT 06117, USADepartment of Pediatrics, University of Minnesota, Minneapolis, MN 55455, USADepartment of Medicine, University of Connecticut Health Center, 263 Farmington Ave, Farmington, CT 06030, USAHyaluronan, a key component of the extracellular matrix, plays a crucial role in joint development and maintenance. In order to determine the role of hyaluronan function in joint development and homeostasis, conditional loss-of-function experiments of Hyaluronan Synthase 2 (<i>Has2</i>) were carried out in mice. <i>Has2</i> depletion in limb mesenchymal cells led to severely shortened limbs with appendicular joints that are deformed, decreased proteoglycan content as characterized by Safranin-O staining, and severely pitted epiphyseal ends of long bones and deformed joints as viewed by micro-CT reconstructions. The embryonic deletion of <i>Has2</i> in mesoderm mesenchyme of limbs by Prx1-Cre confirmed its involvement in joint development, while in situ hybridization and hyaluronan staining confirmed <i>Has2</i> expression and abundant accumulation of hyaluronan in the onset of joint formation, the joint interzone. These findings position <i>Has2</i> as the main hyaluronan-making enzyme in articular cartilage and highlight its essential function in joint formation and retention of proteoglycans of the extracellular matrix of the cartilage.https://www.mdpi.com/2227-9059/13/6/1324synovial joint formationlimb developmentcartilage homeostasishyaluronanhyaluronan synthase 2 (<i>Has2</i>)
spellingShingle Yingcui Li
Alexander Tress
Peter Maye
Kemar Edwards
Asiona Findletar
Nathaniel A. Dyment
Yu Yamaguchi
David W. Rowe
Gengyun Le-Chan
Sunny S. K. Chan
Kevin W.-H. Lo
Genetic Deficiency of Hyaluronan Synthase 2 in the Developing Limb Mesenchyme Impairs Postnatal Synovial Joint Formation
Biomedicines
synovial joint formation
limb development
cartilage homeostasis
hyaluronan
hyaluronan synthase 2 (<i>Has2</i>)
title Genetic Deficiency of Hyaluronan Synthase 2 in the Developing Limb Mesenchyme Impairs Postnatal Synovial Joint Formation
title_full Genetic Deficiency of Hyaluronan Synthase 2 in the Developing Limb Mesenchyme Impairs Postnatal Synovial Joint Formation
title_fullStr Genetic Deficiency of Hyaluronan Synthase 2 in the Developing Limb Mesenchyme Impairs Postnatal Synovial Joint Formation
title_full_unstemmed Genetic Deficiency of Hyaluronan Synthase 2 in the Developing Limb Mesenchyme Impairs Postnatal Synovial Joint Formation
title_short Genetic Deficiency of Hyaluronan Synthase 2 in the Developing Limb Mesenchyme Impairs Postnatal Synovial Joint Formation
title_sort genetic deficiency of hyaluronan synthase 2 in the developing limb mesenchyme impairs postnatal synovial joint formation
topic synovial joint formation
limb development
cartilage homeostasis
hyaluronan
hyaluronan synthase 2 (<i>Has2</i>)
url https://www.mdpi.com/2227-9059/13/6/1324
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