Conformational adaptation of Asian macaque TRIMCyp directs lineage specific antiviral activity.

TRIMCyps are anti-retroviral proteins that have arisen independently in New World and Old World primates. All TRIMCyps comprise a CypA domain fused to the tripartite domains of TRIM5alpha but they have distinct lentiviral specificities, conferring HIV-1 restriction in New World owl monkeys and HIV-2...

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Main Authors: Laura M J Ylinen, Amanda J Price, Jane Rasaiyaah, Stéphane Hué, Nicola J Rose, Flavia Marzetta, Leo C James, Greg J Towers
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2010-08-01
Series:PLoS Pathogens
Online Access:https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1001062&type=printable
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author Laura M J Ylinen
Amanda J Price
Jane Rasaiyaah
Stéphane Hué
Nicola J Rose
Flavia Marzetta
Leo C James
Greg J Towers
author_facet Laura M J Ylinen
Amanda J Price
Jane Rasaiyaah
Stéphane Hué
Nicola J Rose
Flavia Marzetta
Leo C James
Greg J Towers
author_sort Laura M J Ylinen
collection DOAJ
description TRIMCyps are anti-retroviral proteins that have arisen independently in New World and Old World primates. All TRIMCyps comprise a CypA domain fused to the tripartite domains of TRIM5alpha but they have distinct lentiviral specificities, conferring HIV-1 restriction in New World owl monkeys and HIV-2 restriction in Old World rhesus macaques. Here we provide evidence that Asian macaque TRIMCyps have acquired changes that switch restriction specificity between different lentiviral lineages, resulting in species-specific alleles that target different viruses. Structural, thermodynamic and viral restriction analysis suggests that a single mutation in the Cyp domain, R69H, occurred early in macaque TRIMCyp evolution, expanding restriction specificity to the lentiviral lineages found in African green monkeys, sooty mangabeys and chimpanzees. Subsequent mutations have enhanced restriction to particular viruses but at the cost of broad specificity. We reveal how specificity is altered by a scaffold mutation, E143K, that modifies surface electrostatics and propagates conformational changes into the active site. Our results suggest that lentiviruses may have been important pathogens in Asian macaques despite the fact that there are no reported lentiviral infections in current macaque populations.
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spelling doaj-art-0a2cec071f2d4a209356b0a4fedcc6322025-08-20T03:07:20ZengPublic Library of Science (PLoS)PLoS Pathogens1553-73661553-73742010-08-0168e100106210.1371/journal.ppat.1001062Conformational adaptation of Asian macaque TRIMCyp directs lineage specific antiviral activity.Laura M J YlinenAmanda J PriceJane RasaiyaahStéphane HuéNicola J RoseFlavia MarzettaLeo C JamesGreg J TowersTRIMCyps are anti-retroviral proteins that have arisen independently in New World and Old World primates. All TRIMCyps comprise a CypA domain fused to the tripartite domains of TRIM5alpha but they have distinct lentiviral specificities, conferring HIV-1 restriction in New World owl monkeys and HIV-2 restriction in Old World rhesus macaques. Here we provide evidence that Asian macaque TRIMCyps have acquired changes that switch restriction specificity between different lentiviral lineages, resulting in species-specific alleles that target different viruses. Structural, thermodynamic and viral restriction analysis suggests that a single mutation in the Cyp domain, R69H, occurred early in macaque TRIMCyp evolution, expanding restriction specificity to the lentiviral lineages found in African green monkeys, sooty mangabeys and chimpanzees. Subsequent mutations have enhanced restriction to particular viruses but at the cost of broad specificity. We reveal how specificity is altered by a scaffold mutation, E143K, that modifies surface electrostatics and propagates conformational changes into the active site. Our results suggest that lentiviruses may have been important pathogens in Asian macaques despite the fact that there are no reported lentiviral infections in current macaque populations.https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1001062&type=printable
spellingShingle Laura M J Ylinen
Amanda J Price
Jane Rasaiyaah
Stéphane Hué
Nicola J Rose
Flavia Marzetta
Leo C James
Greg J Towers
Conformational adaptation of Asian macaque TRIMCyp directs lineage specific antiviral activity.
PLoS Pathogens
title Conformational adaptation of Asian macaque TRIMCyp directs lineage specific antiviral activity.
title_full Conformational adaptation of Asian macaque TRIMCyp directs lineage specific antiviral activity.
title_fullStr Conformational adaptation of Asian macaque TRIMCyp directs lineage specific antiviral activity.
title_full_unstemmed Conformational adaptation of Asian macaque TRIMCyp directs lineage specific antiviral activity.
title_short Conformational adaptation of Asian macaque TRIMCyp directs lineage specific antiviral activity.
title_sort conformational adaptation of asian macaque trimcyp directs lineage specific antiviral activity
url https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1001062&type=printable
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