Integration of QTL mapping and GWAS reveals the complicated genetic architecture of chemical composition traits in tobacco leaves by Asad Ullah, Asad Ullah, Zhijun Tong, Muhammad Kamran, Muhammad Kamran, Feng Lin, Feng Lin, Tianneng Zhu, Tianneng Zhu, Muhammad Shahzad, Umaira, Umaira, Xuejun Chen, Bingguang Xiao, Haiming Xu, Haiming Xu

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