Chromatin endogenous cleavage provides a global view of yeast RNA polymerase II transcription kinetics

Chromatin immunoprecipitation (ChIP-seq) is the most common approach to observe global binding of proteins to DNA in vivo. The occupancy of transcription factors (TFs) from ChIP-seq agrees well with an alternative method, chromatin endogenous cleavage (ChEC-seq2). However, ChIP-seq and ChEC-seq2 rev...

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Main Authors: Jake VanBelzen, Bennet Sakelaris, Donna G Brickner, Nikita Marcou, Hermann Riecke, Niall M Mangan, Jason H Brickner
Format: Article
Language:English
Published: eLife Sciences Publications Ltd 2024-11-01
Series:eLife
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Online Access:https://elifesciences.org/articles/100764
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author Jake VanBelzen
Bennet Sakelaris
Donna G Brickner
Nikita Marcou
Hermann Riecke
Niall M Mangan
Jason H Brickner
author_facet Jake VanBelzen
Bennet Sakelaris
Donna G Brickner
Nikita Marcou
Hermann Riecke
Niall M Mangan
Jason H Brickner
author_sort Jake VanBelzen
collection DOAJ
description Chromatin immunoprecipitation (ChIP-seq) is the most common approach to observe global binding of proteins to DNA in vivo. The occupancy of transcription factors (TFs) from ChIP-seq agrees well with an alternative method, chromatin endogenous cleavage (ChEC-seq2). However, ChIP-seq and ChEC-seq2 reveal strikingly different patterns of enrichment of yeast RNA polymerase II (RNAPII). We hypothesized that this reflects distinct populations of RNAPII, some of which are captured by ChIP-seq and some of which are captured by ChEC-seq2. RNAPII association with enhancers and promoters - predicted from biochemical studies - is detected well by ChEC-seq2 but not by ChIP-seq. Enhancer/promoter-bound RNAPII correlates with transcription levels and matches predicted occupancy based on published rates of enhancer recruitment, preinitiation assembly, initiation, elongation, and termination. The occupancy from ChEC-seq2 allowed us to develop a stochastic model for global kinetics of RNAPII transcription which captured both the ChEC-seq2 data and changes upon chemical-genetic perturbations to transcription. Finally, RNAPII ChEC-seq2 and kinetic modeling suggests that a mutation in the Gcn4 transcription factor that blocks interaction with the NPC destabilizes promoter-associated RNAPII without altering its recruitment to the enhancer.
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spelling doaj-art-f62bc0e3a4cd428c8956e71647aa94932024-11-28T17:48:43ZengeLife Sciences Publications LtdeLife2050-084X2024-11-011310.7554/eLife.100764Chromatin endogenous cleavage provides a global view of yeast RNA polymerase II transcription kineticsJake VanBelzen0https://orcid.org/0000-0003-0949-3641Bennet Sakelaris1https://orcid.org/0000-0001-8798-584XDonna G Brickner2Nikita Marcou3https://orcid.org/0000-0003-0232-081XHermann Riecke4https://orcid.org/0000-0002-6070-4742Niall M Mangan5https://orcid.org/0000-0002-3491-8341Jason H Brickner6https://orcid.org/0000-0001-8019-3743Department of Molecular Biosciences, Northwestern University, Evanston, United StatesDepartment of Engineering Sciences and Applied Mathematics, Northwestern University, Evanston, United StatesDepartment of Molecular Biosciences, Northwestern University, Evanston, United StatesDepartment of Molecular Biosciences, Northwestern University, Evanston, United StatesDepartment of Engineering Sciences and Applied Mathematics, Northwestern University, Evanston, United StatesDepartment of Engineering Sciences and Applied Mathematics, Northwestern University, Evanston, United StatesDepartment of Molecular Biosciences, Northwestern University, Evanston, United StatesChromatin immunoprecipitation (ChIP-seq) is the most common approach to observe global binding of proteins to DNA in vivo. The occupancy of transcription factors (TFs) from ChIP-seq agrees well with an alternative method, chromatin endogenous cleavage (ChEC-seq2). However, ChIP-seq and ChEC-seq2 reveal strikingly different patterns of enrichment of yeast RNA polymerase II (RNAPII). We hypothesized that this reflects distinct populations of RNAPII, some of which are captured by ChIP-seq and some of which are captured by ChEC-seq2. RNAPII association with enhancers and promoters - predicted from biochemical studies - is detected well by ChEC-seq2 but not by ChIP-seq. Enhancer/promoter-bound RNAPII correlates with transcription levels and matches predicted occupancy based on published rates of enhancer recruitment, preinitiation assembly, initiation, elongation, and termination. The occupancy from ChEC-seq2 allowed us to develop a stochastic model for global kinetics of RNAPII transcription which captured both the ChEC-seq2 data and changes upon chemical-genetic perturbations to transcription. Finally, RNAPII ChEC-seq2 and kinetic modeling suggests that a mutation in the Gcn4 transcription factor that blocks interaction with the NPC destabilizes promoter-associated RNAPII without altering its recruitment to the enhancer.https://elifesciences.org/articles/100764transcriptiongenomicskineticsregulationChIPChEC
spellingShingle Jake VanBelzen
Bennet Sakelaris
Donna G Brickner
Nikita Marcou
Hermann Riecke
Niall M Mangan
Jason H Brickner
Chromatin endogenous cleavage provides a global view of yeast RNA polymerase II transcription kinetics
eLife
transcription
genomics
kinetics
regulation
ChIP
ChEC
title Chromatin endogenous cleavage provides a global view of yeast RNA polymerase II transcription kinetics
title_full Chromatin endogenous cleavage provides a global view of yeast RNA polymerase II transcription kinetics
title_fullStr Chromatin endogenous cleavage provides a global view of yeast RNA polymerase II transcription kinetics
title_full_unstemmed Chromatin endogenous cleavage provides a global view of yeast RNA polymerase II transcription kinetics
title_short Chromatin endogenous cleavage provides a global view of yeast RNA polymerase II transcription kinetics
title_sort chromatin endogenous cleavage provides a global view of yeast rna polymerase ii transcription kinetics
topic transcription
genomics
kinetics
regulation
ChIP
ChEC
url https://elifesciences.org/articles/100764
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AT donnagbrickner chromatinendogenouscleavageprovidesaglobalviewofyeastrnapolymeraseiitranscriptionkinetics
AT nikitamarcou chromatinendogenouscleavageprovidesaglobalviewofyeastrnapolymeraseiitranscriptionkinetics
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