NAC25 transcription factor regulates the degeneration of cytoplasmic membrane integrity and starch biosynthesis in rice endosperm through interacting with MADS29

IntroductionGrain filling is a crucial stage of the rice endosperm development. During this process, the endosperm accumulates abundant storage products such as starch and proteins, which determine both the yield and quality of the grain.MethodsHere, we analyzed the expression of NAC25 transcription...

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Main Authors: Rong Li, Ming-Wei Wu, Jinxin Liu, Xintong Xu, Yiqun Bao, Chun-Ming Liu
Format: Article
Language:English
Published: Frontiers Media S.A. 2025-03-01
Series:Frontiers in Plant Science
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Online Access:https://www.frontiersin.org/articles/10.3389/fpls.2025.1563065/full
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author Rong Li
Rong Li
Ming-Wei Wu
Ming-Wei Wu
Jinxin Liu
Xintong Xu
Yiqun Bao
Chun-Ming Liu
Chun-Ming Liu
Chun-Ming Liu
author_facet Rong Li
Rong Li
Ming-Wei Wu
Ming-Wei Wu
Jinxin Liu
Xintong Xu
Yiqun Bao
Chun-Ming Liu
Chun-Ming Liu
Chun-Ming Liu
author_sort Rong Li
collection DOAJ
description IntroductionGrain filling is a crucial stage of the rice endosperm development. During this process, the endosperm accumulates abundant storage products such as starch and proteins, which determine both the yield and quality of the grain.MethodsHere, we analyzed the expression of NAC25 transcription factor via qRT-PCR and histochemical GUS assays, and obtained its mutants by CRISPR/Cas9-based gene editing in ZH11.Results and discussionThe results showed that NAC25 was expressed specifically in developing rice endosperm, and knockout of NAC25 led to delayed degeneration of cytoplasmic membrane integrity, reduced starch accumulation and chalky starchy endosperm. We showed that NAC25 interacted with MADS29, a MADS family transcription factor whose mutant also showed defective grain filling. These results provide novel insight into the transcriptional regulation of rice grain filling.
format Article
id doaj-art-e9f6dae17ec9449db2ff48b9a32f5f4e
institution DOAJ
issn 1664-462X
language English
publishDate 2025-03-01
publisher Frontiers Media S.A.
record_format Article
series Frontiers in Plant Science
spelling doaj-art-e9f6dae17ec9449db2ff48b9a32f5f4e2025-08-20T03:01:58ZengFrontiers Media S.A.Frontiers in Plant Science1664-462X2025-03-011610.3389/fpls.2025.15630651563065NAC25 transcription factor regulates the degeneration of cytoplasmic membrane integrity and starch biosynthesis in rice endosperm through interacting with MADS29Rong Li0Rong Li1Ming-Wei Wu2Ming-Wei Wu3Jinxin Liu4Xintong Xu5Yiqun Bao6Chun-Ming Liu7Chun-Ming Liu8Chun-Ming Liu9College of Life Sciences, Nanjing Agricultural University, Nanjing, ChinaInstitute of Crop Sciences, Chinese Academy of Agricultural Sciences, Beijing, ChinaKey Laboratory of Plant Molecular Physiology, Institute of Botany, Chinese Academy of Sciences, Beijing, ChinaCollege of Life Sciences, University of Chinese Academy of Sciences, Beijing, ChinaKey Laboratory of Plant Molecular Physiology, Institute of Botany, Chinese Academy of Sciences, Beijing, ChinaSchool of Advanced Agricultural Sciences, Peking University, Beijing, ChinaCollege of Life Sciences, Nanjing Agricultural University, Nanjing, ChinaInstitute of Crop Sciences, Chinese Academy of Agricultural Sciences, Beijing, ChinaKey Laboratory of Plant Molecular Physiology, Institute of Botany, Chinese Academy of Sciences, Beijing, ChinaCollege of Life Sciences, University of Chinese Academy of Sciences, Beijing, ChinaIntroductionGrain filling is a crucial stage of the rice endosperm development. During this process, the endosperm accumulates abundant storage products such as starch and proteins, which determine both the yield and quality of the grain.MethodsHere, we analyzed the expression of NAC25 transcription factor via qRT-PCR and histochemical GUS assays, and obtained its mutants by CRISPR/Cas9-based gene editing in ZH11.Results and discussionThe results showed that NAC25 was expressed specifically in developing rice endosperm, and knockout of NAC25 led to delayed degeneration of cytoplasmic membrane integrity, reduced starch accumulation and chalky starchy endosperm. We showed that NAC25 interacted with MADS29, a MADS family transcription factor whose mutant also showed defective grain filling. These results provide novel insight into the transcriptional regulation of rice grain filling.https://www.frontiersin.org/articles/10.3389/fpls.2025.1563065/fulltranscription factorNAC25MADS29starch synthesiscytoplasmic membrane degenerationrice grain filling
spellingShingle Rong Li
Rong Li
Ming-Wei Wu
Ming-Wei Wu
Jinxin Liu
Xintong Xu
Yiqun Bao
Chun-Ming Liu
Chun-Ming Liu
Chun-Ming Liu
NAC25 transcription factor regulates the degeneration of cytoplasmic membrane integrity and starch biosynthesis in rice endosperm through interacting with MADS29
Frontiers in Plant Science
transcription factor
NAC25
MADS29
starch synthesis
cytoplasmic membrane degeneration
rice grain filling
title NAC25 transcription factor regulates the degeneration of cytoplasmic membrane integrity and starch biosynthesis in rice endosperm through interacting with MADS29
title_full NAC25 transcription factor regulates the degeneration of cytoplasmic membrane integrity and starch biosynthesis in rice endosperm through interacting with MADS29
title_fullStr NAC25 transcription factor regulates the degeneration of cytoplasmic membrane integrity and starch biosynthesis in rice endosperm through interacting with MADS29
title_full_unstemmed NAC25 transcription factor regulates the degeneration of cytoplasmic membrane integrity and starch biosynthesis in rice endosperm through interacting with MADS29
title_short NAC25 transcription factor regulates the degeneration of cytoplasmic membrane integrity and starch biosynthesis in rice endosperm through interacting with MADS29
title_sort nac25 transcription factor regulates the degeneration of cytoplasmic membrane integrity and starch biosynthesis in rice endosperm through interacting with mads29
topic transcription factor
NAC25
MADS29
starch synthesis
cytoplasmic membrane degeneration
rice grain filling
url https://www.frontiersin.org/articles/10.3389/fpls.2025.1563065/full
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